Quedius (Raphirus) tergimpressus, Smetana, Aleš, 2012

Smetana, Aleš, 2012, Contributions to the knowledge of the Quediina (Coleoptera, Staphylinidae, Staphylinini) of China. Part 42. Genus Quedius Stephens, 1829. Subgenus Raphirus Stephens, 1829. Section 10, Zootaxa 3156, pp. 43-68 : 57-60

Quedius (Raphirus) tergimpressus sp. nov.

( Figs. 54–61 )

Type locality. People’s Republic of China: Yunnan: Baoshan Pref., Gaoligong Shan, 31 km SE Tengchong, 24°53ʹ11ʺ N 98°45ʹ22ʺE, 2200–2250 m.

Type material. Holotype (male) and allotype (female): China: CHINA (Yunnan) Baoshan Pref., Gaoligong Shan 31 km SE Tengchong 2200–2250 m 24°53ʹ11ʺ N / 98°45ʹ22ʺE (pasture in second. forest, under moss/shrubs) 26.VIII.2009 D. W. Wrase [09] . Holotype in MSC, allotype in ASC.

Paratypes: China: Yunnan: same data as holotype, 13, 1♀ ( ASC, MSC) ; Baoshan Pref., Gaoligong Shan E pass, 36 km SE Tengchong, 2200 m (farm land under moss, grass, shrubs, under stones) 24°49ʹ32ʺN / 98°46ʹ0 6ʺE 28.VIII. 2009, D. E. Wrase [13], 2♀ ( MSC) ; Lincang Pref., Bangma Shan, E pass, 17 km NW Lincang 2040 m (secondary pine forest with ferns, litter sifted) 23°57ʹ31ʺN / 99°56ʹ13ʺE 9.IX. 2009, D. W. Wrase [36], 13 ( MSC) ; Lincang Pref., Xue Shan, 48 km N Lincang 2070 m 2419ʹ0 3ʺN 10007ʹ13ʺE (slope close to small pond, in loamy soil/plant roots) 12.IX. 2009, D. W. Wrase [45], 13 ( ASC).

Description. In all external characters similar to Q. aureiventris Bernhauer, 1915 (detailed description in Smetana 1988: 250), and different mainly by the male sexual characters, particularly by the shape of the aedoeagus and especially by that of the paramere, by the less dilated first four segments of the male front tarsus, and by the uniformly dark pubescence of the elytra (elytral pubescence in Q. aureiventris is a mixture of dark and golden or golden-yellowish hairs).

M a l e. First four segments of front tarsus slightly dilated, sub-bilobed, each with tenent setae ventrally, segment 2 slightly narrower than apex of tibia (ratio 0.88), segment 4 narrower than preceding segments. Sternite 8 with two long setae on each side, with deep and wide medioapical emargination, small triangular area before emargination flattened and smooth. Genital segment with tergite 10 slightly wider and less setose than that of Q. aureiventris ( Fig. 51 , 55 ); sternite 9 with basal portion short and wide, apical portion with apex narrowly arcuate, with differentiated pair of apical setae, general setae long ( Fig.56 ). Aedoeagus ( Figs. 57–60 ) narrow, elongate, median lobe subparallelsided, anteriorly not widened before narrowed into long apical portion with subacute apex, on face adjacent to paramere, when paramere removed, with short median carina forming a distinct dent in lateral view ( Figs. 58, 59 ); paramere markedly different from those of both Q. aureiventris and Q. impressiventris , distinctly fusiform, with narrowly arcuate apex not quite reaching apex of median lobe; four fine setae at apical margin, medial setae longer than lateral ones, two similar setae at each lateral margin below apex; sensory peg setae on underside of paramere numerous, forming one long, fairly regular row along each margin of apical half of paramere ( Fig. 60 ).

F e m a l e. First four segments of front tarsus not appreciably dilated. Tergite 10 of genital segment shorter and wider than that of Q. aureiventris and less setose in front of long apical setae ( Figs. 53 , 61 ).

Length 4.7–5.0 mm.

Geographical distribution. Quedius tergimpressus is at present known from several localities in Gaoligong Shan, Bangma Shan and Xue Shan, all in western Yunnan.

Bionomics. Specimens of the original series were taken in farmland from under moss and stones, and among grass; in a secondary forest by sifting litter among ferns, and on a slope close to small pond in loamy soil with plant roots.

Recognition and comments. Quedius tergimpressus differs from Q. aureiventris by the characters given in the description and the differences against Q. impressiventris are given in Comments under the latter species. In addition, Q. tergimpressus differs from both Q. aureiventris and Q. impressiventris by the less dilated first four segments of the male front tarsus.

Etymology. The specific epithet is a combination of the Latin noun tergum and the adjective impressus, - a, um (impressed). It refers to the impressions on the abdominal tergites of the species.

Figures

FIGURES 43 – 54. 43 – 50. Quedius impressiventris: 43, tergite 10 of male genital segment; 44, sternite 9 of male genital segment; 45, aedoaegus, ventral view; 46, apical portion of median lobe, lateral view, paramere removed; 47, apical portion of median lobe, ventral view, paramere removed; 48, 49, apical portions of undersides of parameres with sensory peg setae; 50, tergite 10 of female genital segment. 51 – 53. Quedius aureiventris: 51, tergite 10 of male geital segment; 52 sternite 9 of male genital segment; 53, tergite 10 of female genital segment. Quedius tergimpressus: 54, apical portion of male sternite 8.

FIGURES 55 – 65. 55 – 61. Quedius tergimpressus: 55, tergite 10 of male genital segment; 56, sternite 9 of male genital segment; 57, aedoeagus, ventral view; 58, apical portion of median lobe, ventral view, paramere removed; 59, apical portion of median lobe, lateral view, paramere removed; 60, apical portion of underside of paramere with sensory peg setae; 61, tergite 10 of female genital segment. 62 – 65. Quedius meilixue: 62, apical portion of male sternite 8; 63, tergite 10 of male genital segment; 64, sternite 9 of male genital segment; 65, aedoeagus, ventral view.