Cambalida griswoldi, Haddad, Charles Richard, 2012

Haddad, Charles Richard, 2012, A revision of the Afrotropical spider genus Cambalida Simon, 1909 (Araneae, Corinnidae), ZooKeys 234, pp. 67-119 : 100-102

Cambalida griswoldi sp. n. Figures 55, 84-86

Type material.

Holotype female. MADAGASCAR: Antsiranana : Réserve Spéciale d’Ambre, 3.5km 235°SW Sakaramy, 12°28'08"S, 49°14'32"E, 325m a.s.l., leg. Fisher-Griswold Arthropod Team, 26-31.I.2001 (sifted litter, tropical dry forest) (CAS, CASENT 9006738).

Paratypes.

MADAGASCAR: Antsiranana : Forêt d’Orangea, 3.6km 128° SE Remena, 12°15'32"S, 49°22'29"E, 90m a.s.l., leg. Fisher, Griswold et al., 22-28.II.2001 (pitfall trap, littoral rainforest), 2♀ (CAS, CASENT 9007088); Same locality as holotype, leg. L.J. Boutin, 26-31.I.2001, 1♀ (CAS, CASENT 9000791). Toliara : 18km NNW Betroka, 23°09'48"S, 45°58'07"E, 825m a.s.l., leg. M. Ivie & A. Pollock, 9-14.XII.1994 (flight intercept traps), 1♂ (CAS, CASENT 9033123).

Other material examined.

None.

Diagnosis.

Females have similar spermathecal proportions to the continental Cambalida fulvipes but can be recognised by the narrower, distinctly coiled epigynal ridges (Fig. 84), which are broad and curved in Cambalida fulvipes (Fig. 80). The distal section of the male embolus is short and slightly curved towards the tip (Fig. 86).

Etymology.

A patronym in honour of Charles Griswold, one of the collectors of the types, in recognition of his vast and significant collections of spiders on Madagascar.

Remark.

The palp of the paratype male is greatly expanded and only the tegulum is illustrated here (Fig. 86). The cymbium has the distal setae characteristic of the genus, confirming the placement of this species and the occurrence of Cambalida on Madagascar.

Female

(holotype, Réserve Spéciale d’Ambre, CASENT 9006738). Measurements: CL 2.10, CW 1.52, AL 2.53, AW 1.78, TL 4.70 (4.65-4.75), FL 0.13, SL 0.97, SW 0.89, AME–AME 0.05, AME–ALE 0.02, ALE–ALE 0.26, PME–PME 0.11, PME–PLE 0.06, PLE–PLE 0.43, PERW 0.62, MOQAW 0.23, MOQPW 0.37, MOQL 0.36.

Length of leg segments (sequence from femur to tarsus, and total): I 1.28 + 0.57 + 1.03 + 0.98 + 0.64 = 4.50; II 1.17 + 0.55 + 0.84 + 0.88 + 0.56 = 4.00; III 1.05 + 0.54 + 0.70 + 0.95 + 0.44 = 3.68; IV 1.48 + 0.61 + 1.28 + 1.57 + 0.58 = 5.52.

Carapace deep orange-brown, clypeus deep yellow-brown, eye region slightly darker; faint black striae radiating from fovea towards palps and coxae and faint black mottling on slopes; surface finely granulate, sparsely covered in white plumose setae. AER procurved, eyes subequal in diameter; AME separated by distance ½ their diameter, AME separated from ALE by distance equal to ¼ AME diameter; clypeus height equal to 1⅔ AME diameter; PER procurved, PLE slightly larger than PME; PME separated by distance slightly more than 4∕5 their diameter, PME separated from PLE by distance slightly less than ½ PME diameter; CW:PERW = 2.45:1. Chelicerae deep yellow-brown with black mottling on anterior surface; three teeth on promargin, median tooth largest, proximal and distal teeth small, subequal in size, median and distal teeth adjacent; two teeth on retromargin, distal tooth slightly larger than proximal tooth, closer to fang base than promarginal teeth; endites yellow with black mottling, cream prolaterally; labium orange-brown with faint black mottling, cream distally; sternum orange with dense brown mottling, except at setal bases. Legs finely granulate; femora I–IV yellow-brown with black mottling laterally and ventrally, absent dorsally and at distal end ventrally; patellae I–IV yellow-brown with black mottling laterally and ventrally, absent dorsally, with faint median dorsal black line; tibiae I yellow with black mottling in distal half; tibiae II and III yellow-brown and IV orange-brown, all with black mottling, faint dorsally, with fine median dorsal black line; metatarsi I–III yellow with black mottling, absent proximally; metatarsi IV yellow-brown with black mottling, faint dorsally, absent proximally and distally; tarsi I–IV yellow. Leg spination: femora: I pl 1 do 3, II do 1, III pl 1 do 2 rl 1, IV pl 1 do 3 rl 1; patellae with do 1 long distal seta; tibiae: I plv 3 rlv 2, II rlv 2, III pl 2 rl 2 plv 2 rlv 1, IV pl 2 rl 2 plv 2 rlv 1 vt 2; metatarsi: I plv 2 rlv 2, II plv 2 rlv 2, III pl 2 rl 2 plv 2 rlv 2 vt 3, IV pl 3 rl 3 plv 2 rlv 2 vt 3. Palpal spination: femora do 2, patellae pl 1, tibiae pl 1 plv 1, tarsi pl 1 plv 3. Abdomen mottled dark grey dorsally with fine cream chevrons posteriorly; dorsum with red-brown scutum extending ⅔ abdomen length, with black mottling darker around margins of scutum, forming ring-shaped dorsal marking; venter mottled pale grey, darker towards spinnerets, with paired rectangular black markings between paired sclerite rows, midway between epigastric furrow and spinnerets; epigastric sclerite red-brown and inframamillary sclerite yellow-brown. Epigyne with small lateral copulatory openings situated within 6-shaped epigynal ridges (Fig. 84); copulatory ducts initially directed transversely medially, bending at nearly right angle, entering ST II posterolaterally; ST II round, with nearly right-angled posterolateral margin, joined broadly to kidney-shaped posterior ST I (Fig. 85).

Male

(paratype, Betroka, CASENT 9033123). Measurements: CL 2.23, CW 1.72, AL 2.70, AW 1.61, TL 4.83, FL 0.15, SL 1.03, SW 0.94, AME–AME 0.06, AME–ALE 0.02, ALE–ALE 0.30, PME–PME 0.13, PME–PLE 0.06, PLE–PLE 0.48, PERW 0.71, MOQAW 0.29, MOQPW 0.41, MOQL 0.44.

Length of leg segments (sequence from femur to tarsus, and total): I 1.53 + 0.63 + 1.30 + 1.24 + 0.95 = 5.65; II 1.38 + 0.60 + 1.05 + 1.08 + 0.81 = 4.92; III 1.23 + 0.57 + 0.90 + 1.11 + 0.65 = 4.46; IV 1.89 + 0.74 + 1.69 + 1.90 + 0.83 = 7.05.

Carapace dark brown, eye region darker; faint black striae radiating from fovea towards palps and leg coxae; surface finely granulate, densely covered in white plumose setae. Rings around eyes faded to orange-brown; AER procurved, ALE larger than AME; AME separated by distance equal to ½ their diameter, AME separated from ALE by 1∕5 AME diameter; clypeus height slightly less than 1½ AME diameter; PER procurved, PLE slightly larger than PME; PME separated by distance slightly less than ¾ their diameter, PME separated from PLE by distance slightly less than ⅓ PME diameter; CW:PERW = 2.42:1. Chelicerae brown with faint black mottling on anterior surface, yellow prolaterally in distal half; three teeth on promargin, median tooth largest, distal tooth smallest, median and distal teeth closest; two slightly separated teeth on retromargin, distal tooth slightly smaller than proximal tooth, closer to fang base than promarginal teeth; endites pale brown with dark brown mottling, cream prolaterally; labium pale brown with dark brown mottling, cream distally; sternum red-brown with dark brown mottling. Legs finely granulate; femora I–IV dark brown, slightly paler dorsally, yellow dorsally at distal end; patellae I–IV yellow with black mottling laterally and ventrally distally, darker on posterior legs; tibiae I and II yellow with black mottling; tibiae III and IV orange-brown with dense black mottling, paler along dorsal midline; metatarsi I and II yellow-brown, with sparse dark brown mottling laterally; metatarsi III and IV dark brown, yellow at proximal and distal ends; tarsi I–IV yellow, tarsus IV with black mottling medially. Leg spination: femora: I pl 1 do 3, II do 3, III pl 1 do 3 rl 1, IV pl 1 do 3 rl 1; patellae with do 1 long distal seta; tibiae: I and II spineless, III pl 2 rl 2 plv 2, IV pl 2 rl 2 plv 2 vt 2; metatarsi: I plv 2 rlv 1, II plv 1 rlv 1, III pl 2 rl 1 plv 1 vt 3, IV pl 3 rl 3 plv 2 vt 3. Palpal spination: femora pl 1 do 2, patellae pl 1, tibiae pl 1 plv 1, tarsi pl 1 plv 3. Abdomen with dark red-brown, nearly black scutum covering entire dorsum; venter mottled pale grey between sclerites; epigastric sclerite, post-epigastic sclerites and ventral sclerite deep red-brown with black mottling, inframamillary sclerite orange-brown. Palps orange-brown with dense black mottling; embolus short, basal coil slightly curved, distal section gently curved towards tip (Fig. 86).

Distribution.

Known from isolated localities in northern and southern Madagascar (Fig. 95).

Biology.

A ground-dwelling species collected in contrasting forest types.