Victoriana melanorrhina (Philippi, 1865) Miranda & Skevington & Marshall, 2020

Miranda, Gil Felipe Gonçalves, Skevington, Jeffrey H. & Marshall, Stephen A., 2020, New generic concepts for orphaned lineages formerly treated as part of the genus Ocyptamus Macquart, 1834 (Diptera, Syrphidae), Zootaxa 4822 (2), pp. 151-174 : 163

Victoriana melanorrhina species group (Victoriana sensu stricto)

Description. Body brown. Head. Face narrow to normal (between 1/4 to ~1/3 of head width), entirely pale or with medial dark stripe; tubercle dorsally positioned. Antennal insertions confluent. Frons ~1/3 of head width. Female ocellar triangle ~1 ocellus-width from lateral eye margin. Dorsal occiput with one row of pile; *ventral occiput with rows of pile distanced from eye margin ( Fig. 88 ). Thorax. Scutum dark, sometimes pale laterally, and without distinct anterior row of pile. Scutellum pale to mostly dark; *sub-scutellar fringe conspicuous, but pile sparse. Anterior anepisternum pilose. Katatergite with short microtrichia that gives the sclerite a ‘velvet’ appearance. Metaepisternum pilose. Metasternum bare. Upper calypter margin with shorter to much shorter pile than pile on the ventral calypter margin. *Metafemur with erect pile on its baso-posterior surface ( Fig. 90 ). Wing. Alula absent, or much reduced (narrower than c) ( Fig. 91 ). Wing hyaline or with anterior dark margin on females, light brown on males; vein M1 strongly oblique, ending far from where it leaves M ( Fig. 48 ). Abdomen. Abdomen parallel-sided, elongated, slightly petiolate; *terga 4 and 5 with sub-basal, pale, small, banded maculae ( Fig. 48 ). Terminalia. Female tergum 7 as pair of sclerotized stripes ( Fig. 100 ); tergum 8 as a narrow triangular sclerotization ( Fig. 100 ). *Male postgonite elongated, with a dorsal acute extremity and a ventral acute extremity (that may be further extended anteriorly) ( Figs 54 and 55 ); *surstylus elongated and finger-like, ventral surface with setae and setulae intermixed ( Figs 51 and 52 ); basiphallus teardrop-shaped, distiphallus membranous with dorsal sclerotized triangular region (as in Fig. 97 ).

Included species (3). V. laudabilis ( Williston, 1891) comb. nov. [1b], V. lugubris ( Philippi, 1865) comb. nov. [2, 3], V. melanorrhina ( Philippi, 1865) comb. nov. [2, 3, 4].

Figures

FIGURES 78–88. 78: Fragosa harlequina (Hull) (holotype Baccha harlequina Hull, AMNH), male, thorax, lateral, by AMNH (line drawing represents outline of the alula). 79: Fragosa argentina (Curran) (CNC_Diptera174232), thorax, postero-ventral (line drawing represents outline of the incomplete post-metacoxal bridge). 80–81: Fragosa virgilio (Hull) (holotype Baccha virgilio Hull, CNC, CNC_Diptera189124), male. 80: habitus, lateral, by CNC. 81: head, oblique frontal, by CNC. 82: Fragosa titania (Hull) (JSS25229), female, head, dorsal. 83: Fragosa stenogaster (Williston) (debu01088840), male, head, dorsal. 84–85: Hypocritanus fascipennis (Wiedemann), female, head. 84: frontal, by Ximo Mengual (http://syrphidae.myspecies.info/ taxonomy/term/879). 85: lateral (CNC_Diptera501). 86–87: Nuntianus cecrops (Hull) (INPA-DIP000246), female, head. 86: dorsal. 87: lateral. 88: Victoriana aff. melanorrhina (DEBU), female, head, lateral. a: alula; ai: antennal insertion; f: frons; ft: frontal triangle; lem: lateral eye margin; mt: metafemur; o: ocellar triangle; oc: occiput; pmt: post-metacoxal bridge; t: facial tubercle.

FIGURES 89–102. 89: Victoriana sativa (Curran) (holotype Baccha sativa Curran, AMNH), habitus, dorsal, by AMNH. 90– 91: Victoriana laudabilis (Williston) (holotype Baccha laudabilis Williston, BMNH, NHMUK010369870), copyright of The Trustees of the Natural History Museum, London. 90: base of abdomen and metafemur, dorsal. 91: base of wing, dorsal (line drawing represents outline of the alula). 92–95: Fragosa sp. (species group tenuis, CNC_Diptera196012), male. 92: hypandrium, lateral. 93: surstylus, ventro-lateral. 94: epandrium, lateral. 95: epandrium, ventral (line drawing represents outline of the subepandrial sclerite). 96–97: Nuntianus abata (Curran) (CNC_Diptera209361), male, hypandrium. 96: lateral. 97: ventral. 98: Syrphus conjunctus Wiedemann (currently incertae sedis, CNC_Diptera209347), male, hypandrium, lateral. 99: Victoriana aff. melanorrhina (debu00279441), male, hypandrium, ventral. 100: Victoriana aff. melanorrhina (DEBU), female, terminalia, dorsal. 101: Victoriana parvicornis (Loew) (CNC_Diptera237922), terminalia, dorsal. 102: Maiana pumila (Austen) (debu00178088), female, terminalia, dorsal. a: alula; c: cercus; d: distiphallus; e: epiproct; l: lobe; p: baso-posterior pile; t7 and 8: tergum 7 and 8.

FIGURES 48–60. Victoriana Miranda gen. nov.. 48: Victoriana melanorrhina (Philippi), female, habitus, dorsal, by Steve Marshall, photo previously published in Mengual et al. (2018). 49: Victoriana parvicornis (Loew) (CNC_Diptera237923), male, habitus, dorsal, photo previously published in Mengual et al. (2018). 50: Victoriana zilla (Hull) (holotype Baccha zilla Hull, AMNH), female, habitus, dorsal, by AMNH. 51: Victoriana aff. melanorrhina (debu00279441), male, epandrium, lateral. 52: Victoriana aff. melanorrhina (debu00279441), male, epandrium, ventral. 53: Victoriana cf. attenuata (JSS22253), male, epandrium, ventral. 54: Victoriana aff. melanorrhina (debu00279441), male, hypandrium, lateral. 55: Victoriana melanorrhina (debu01088955), male, hypandrium, lateral. 56: Victoriana cf. attenuata (JSS22253), male, hypandrium, lateral. 57–60. Victoriana parvicornis (Loew) (CNC_Diptera237923), male. 57: abdomen apex, ventral. 58: epandrium, frontal. 59: hypandrium, ventral. 60: hypandrium, lateral. c: cercus; d: distiphallus; e: epandrium; ex: extensions of sternum 5; p: postgonite; pr: process of sternum 8; s: surstylus; se: subepandrial sclerite; s8: sternum 8.