Artoria quadrata Framenau, 2002

Framenau, Volker W. & Baehr, Barbara C., 2018, The wolf spider genus Artoria in New South Wales and the Australian Capital Territory, Australia (Araneae, Lycosidae, Artoriinae), Evolutionary Systematics 2 (2), pp. 169-241 : 169

Artoria quadrata Framenau, 2002 Figs 34, 35 A–H, 46C Square Forest Runner

Artoria quadrata Framenau, 2002: 224-226, figs 19 A–F, 20.

Material examined.

Holotype male, Avon River near Valencia Creek (37°48'S 146°27'E, Victoria, AUSTRALIA], 18 December 1996, riparian gravel bank, V.W. Framenau (NMV K7678). Paratypes: 3 females, same data as holotype, except 18 December 1996-1 January 1997, pitfall trap (MV K7679-80).

Other material examined.

152 males, 88 females (3 with eggsac) and 30 juveniles in 91 records (85 NSW, 6 ACT). AUSTRALIA: Australian Capital Territory: 1 male, Black Mountain, 35°16'S, 149°06'E (QM S64055); 1 male, Black Mountain, W slope (Caswell Drive), 35°16'S, 149°06'E (ANIC); 1 male, 1 female, Canberra, 35°18'S, 149°08'E (NMV K7758); 1 female, Mt Ainslie, NW face, 35°16'S, 149°10'E (ANIC); 3 females, 1 juv., Tidbinbilla Nature Reserve, 35°28'S, 148°52'E (AM KS13964); 1 male, Yarralumla, 35°19'S, 149°04'E (ANIC). New South Wales: 1 male, `Temi` (N of Murrurundi), Chilcotts Creek Road, 31°40 ’48” S, 150°49 ’00” E (AM KS82850); 1 female, `Temi`, off Chilcott’s Creek Road, near gate, off New England Highway, approx. 7 km S of Willow Tree, 31°40 ’48” S, 150°49 ’01” E (AM KS75586); 1 female, 3 km east of New England Highway and 3 km east of Ardglen, 31°43 ’44” S, 150°49 ’10” E (AM KS123042); 1 female, 400 m from Moss Vale Road and 5 km NW of Moonan Flat, private land, 31°54 ’13” S, 151°11 ’54” E (AM KS123083); 1 female, 500 m off Barrington West Road and 3 km S of Barrington, private land, 31°59 ’42” S, 151°54 ’08” E (AM KS122814); 4 females, 70 m from Coolah Creek Road and 11 km northeast of Coolah, private land 'Mt Mill’, 31°46 ’20” S, 149°49 ’02” E (AM KS123019); 2 males, 4 females, 80 m from Cassilis Road and 20 km NW of Cassilis, private land, 31°54 ’01” S, 149°48 ’21” E (AM KS122586, KS123016); 5 males, 4 females, 9 juv., Abercrombie Caves area, near Bushrangers Cave, 33°54 ’45” S, 149°21 ’28” E (AM KS22494); 1 male, 5 juv., Abercrombie Caves area, through arch near Bushrangers Cave, 33°54'S, 149°22'E (AM KS21686); 1 female, Abercrombie Caves, entrance arch, 33°54 ’45” S, 149°21 ’28” E (AM KS5523); 33 males, 9 females, 12 juv., Abercrombie Caves, near Bushrangers Cave, 33°54'S, 149°22'E (AM KS22469, KS22486); 3 females, approx. 10 m from Creek, 4 km SW of Mt Vincent, private land, 32°55 ’41” S, 151°26 ’30” E (AM KS122336); 4 females, Attunga State Forest, S of Ardey Range, W edge of SF, opposite `Tralee`, 30°56 ’00” S, 150°54 ’19” E (AM KS82848, KS82861); 1 female, Bellinger River Station, about 600 m down small track off Dorrigo-Bellingen Highway 1 km W of Summervilles Road, 30°25 ’03” S, 152°45 ’30” E (AM KS85360); 2 males, Bullaburra Red Gum Park, 33°44'S, 150°25'E (AM KS51841); 1 male, Bungendore NW, Mt Fairy, 35°9'S, 149°35'E (QM S21907); 2 males, Bungonia, 34°48'S, 150°01'E (WAM T55468); 1 male, Bungonia Caves Area, near Information Centre, 34°48'S, 150°01'E (AM KS71666); 1 male, 1 female, Christophersons Mountain, 2 km SSW of Bostobrick, 30°17 ’45” S, 152°37 ’04” E (AM KS61739); 1 female, Coolah Tops National Park, 200 m from Warung State Forest Road, 31°44 ’01” S, 150°02 ’17” E (AM KS123028); 16 males, 3 females, Coolah Tops National Park, The Forest Road, 0.3 km E of Hildergard Road, 31°43 ’59” S, 150°02 ’04” E (AM KS75425); 2 females, Coonabarabran, 'Smoky Hollow`, 31°16'S, 149°17'E (AM KS7555, KS7558); 2 males, 3 females, Crown Reserve, Barraba-Bundarra Road, N of `Ironbark`, 30°19 ’46” S, 150°46 ’34” E (AM KS82847); 1 female, Dorrigo National Park, 30°22'S, 152°45'E (ANIC); 1 female, Dorrigo National Park, Wonga Walk, 30°22'S, 152°45'E (ANIC); 5 females, 3 juv., Dorrigo National Park, Wonga Walk, 200 m SW of Hardwood Lookout, 30°22 ’53” S, 152°44 ’00” E (AM KS35660); 3 females, Doubleduke State Forest, 1 km SW of junction of Pacific Highway and Glencoe Road, 29°12 ’46” S, 153°15 ’29” E (AM KS39701); 1 male, Ferntree Gully Reserve, approx. 170 m from Bylong Valley Way and 4 km SW of Ginghi, 32°37 ’20” S, 150°03 ’37” E (AM KS128880); 21 male, 1 femal, Hazelbrook, Winbourne Road, 33°43 ’20” S, 150°27 ’25” E (AM KS52081, KS52085-6, KS52088, KS53941, KS53942-3, KS53946-7, KS53948, KS53950-2); 1 female, Illunie Nature Reserve, 34°08 ’33” S, 148°37 ’37” E (AM KS117678); 1 female, Koorawatha Nature Reserve, 34°02 ’31” S, 148°34 ’35” E (AM KS128878); 1 male, Linton Nature Reserve, SW corner of Reserve, 60 m E of road, 30°27 ’45” S, 150°51 ’46” E (AM KS82859); 1 male, Marsfield, Sydney, 33°45'S, 151°06'E (WAM T42123); 1 female, Mororo State Forest, 29°19 ’06” S, 153°14 ’28” E (AM KS88496); 1 male, 1 female, Mt Kaputar National Park, 250 m S of track to car park at Waa Gorge, 30°03 ’59” S, 150°05 ’29” E (AM KS82851KS82856); 1 female, Mt Kaputar National Park, base of W face of Mt Waa, 30°03 ’59” S, 150°06 ’05” E (AM KS82855); 2 males, Mt Kaputar National Park, end of Bullawa Creek Road, 50 m NE of picnic area, 30°14 ’04” S, 150°06 ’39” E (AM KS82857); 1 female, Nadgee Nature Reserve, 37°22'S, 149°55'E (AM KS86431); 1 female, New England National Park, 30°29'S, 152°30'E (QM S64062); 2 females, Nullica Beach, 37°06'S, 149°53'E (AM KS83719); 1 female, Orara State Forest, junction of Urumbilum Creek Road and Loudens Road, 30°19 ’35” S, 152°55 ’00” E (AM KS61026); 8 males, Oxley Wild River National Park, East Kunderang Road, 30°49'S, 152°07'E (AM KS124710KS124746); 18 males, Oxley Wild Rivers National Park, East Kunderang Track, 30°49'S, 152°07'E (AM KS124222); 2 females, Oxley Wild Rivers National Park, Yarrowitch River, 31°04 ’33” S, 152°03 ’23” E (AM KS 124544, KS125238); 1 male, Pacific Palms, 32°21'S, 152°31'E (AM KS85363); 1 female, Ponderosa Forest Park, Nundle State Forest, 31°27 ’30” S, 151°15 ’00” E (QM S64064); 1 female, Ramornie State Forest, Main Creek, track off Mt Tindal Road, 29°43 ’01” S, 152°38 ’24” E (AM KS39706); 1 female, Schofields, 33°42'S, 150°52'E (AM KS85365); 1 female, Severn State Forest, Atholwood Loop Road, 29°04 ’28” S, 151°00 ’53” E (AM KS82849); 1 female, The Battery Picnic Area, SE of Merriwa, 33°12 ’46” S, 150°27 ’31” E (AM KS75037); 2 males, 3 females, Upper Hunter River, Kayuga Bridge at Muswellbrook., 32°15 ’12” S, 150°53 ’16” E (AM KS91854); 1 male, 2 females. Warrumbungle National Park, Camp Pincham Carpark, 31°16'S, 148°55'E (AM KS75177, KS75188, KS75190); 1 female, Wombayan Caves area, 34°19'S, 149°59'E (AM KS85364); 7 males, Wombayan Caves area, hillside north of camping ground, 34°18'S, 149°58'E (AM KS71665, KS85362); 1 male, Wombeyan Caves area, N of campground, 34°19'S, 149°59'E (AM KS85361); 16 males, 1 female, Yabbra Scrub, Yabbra State Forest, 28°38'S, 152°30'E (AM KS51287, KS57769, KS53795, KS63741, KS63745); 1 female, Yengo National Park, approx 40m from track, 32°56 ’46” S, 150°54 ’33” E (AM KS128879).

Diagnosis.

Artoria quadrata is part of the lineata -group, to which also A. lineata and A. ulrichi belong. A detailed diagnosis for these three species can be found above under the treatment of A. lineata .

Description.

Artoria quadrata has been described in detail ( Framenau 2002). A diagnosis and diagnostic images (Figs 35 A–H, 46C) are provided here to facilitate identification.

Life history and habitat preferences.

Artoria quadrata appears to be versatile with respect to its habitat preferences, reported from a variety of environments including woodland and forest habitats (e.g. 'undisturbed forest’, 'scattered trees’, 'semi-cleared woodland adjacent to pine plantations’, 'temperate rainforest’ ' Nothofagus rainforest’, 'dry subtropical rainforest’), and also open habitats - some near water - ('open grassy field, swampy’, 'open grassy field, edge of river’, 'in grass’, ‘garden’, 'bare limestone outcrop’). In NSW and the ACT mature spiders have been found all year round, with a peak from October to January. Three females with eggsacs were found in November.

Distribution.

Artoria quadrata is common east and west of the Great Dividing Range, from coastal areas into the Brigalow Belt South (BBS) and NSW South Western Slopes (NSS) IBRA regions (Fig. 34). The species otherwise occurs in Queensland and Victoria ( Framenau 2002; 2005).