Actenodia Laporte de Castelnau, 1840

Bologna, M. A. & Pinto, J. D., 2002, The Old World genera of Meloidae (Coleoptera): a key and synopsis, Journal of Natural History 36 (17), pp. 2013-2102 : 2061-2062

42. Actenodia Laporte de Castelnau, 1840

(gure 11)

Rusadiria Pardo Alcaide, 1954 (as a subgenus of Mylabris ). Type species: Mylabris billbergi

Gyllenhal, 1817, by original designation. Pardo Alcaide originally introduced this name in 1952 but failed to designate a type species.

2014 Notes Type). 20 species spp.. Actenodia guttata Laporte de Castelnau, 1840 , by monotypy (see February Madagascar Geographic). Records distribution from. Pakistan Mediterranean and Afghanistan Basin, Arabia are questionable and Africa. (except 13 References 15 Taxonomy. Reiche (1865); Marseul (1870, 1872 keys and catalogues); Baudi di: 23 Selve (1878), Wellman (1908 pars); Péringuey (1909); Soumacov (1915); Pardo at Alcaide (1952, 1954a, 1959, 1963); Kuzin (1954); Kaszab (1958b); Bologna (1978, Bath] 1990 Bionomics, 1991a);. Dvorak Gorriz (1993 y Muñoz a). (1882); Beauregard (1890); Bedel (1892); Cros of (1930); Valladares and Salgado (1983); Bologna (1991a). University Larvae Anatomy Pharmacology.. Cros Beauregard. (1917 Beauregard a, 1930 (1890); () 1890 Bologna; Gupta). ((1971 1991, a 1978).). [by Notes Downloaded the Mylabris synonym Selander type and (of Gorrizia Meloe (1991 showed))] decemguttata. indicated It that also decemguttata is, the erroneously Thunberg type species is,, 1791 a that distinct of. the Pardo Arithmema type species Alcaide species and of (, Hycleus 1958 consequently A. b guttata ) examined [cited, is the as a latter name is a junior synonym of Hycleus (see below) and not a senior synonym of Actenodia as listed by Selander (1991).

As discussed by Bologna (1978, 1991a, 2000a), the number of antennal segments is variable in mylabrine genera. For example, in Actenodia the number varies from six to nine (most commonly eight or nine). This variation has caused considerable confusion in generic assignments. For this reason, the number of described species assignable to Actenodia is unclear. Bologna (1991a: 46) noted 31 species (indicated erroneously as ca 50 on p. 275), but it now appears that certain SW African species with eight antennal segments (amoenus Marseul,1872; deserticolus Wellman, 1908

(= annulipes Pic, 1910); bushmanicus Kaszab, 1952; devylderi Borchmann, 1928; kochi

Kaszab, 1952; politus Kaszab, 1955 ; vansoni Kaszab, 1952) should be placed in

Hycleus (Bologna, unpublished). The placement in Actenodia of other African and

Asian species (mirabilis Kaszab, 1952; waziristanica Kaszab, 1958) that we have not examined, also remains questionable.

Dvorak (1993a) recently described Paractenodia perfuga from Syria. This very

2014 February 13 15: 23 at] Bath of University [by Downloaded

2062 M. A. Bologna and J. D. Pinto distinct species has unique coloration (a red frontal spot and red pronotum), only seven antennal segments (abruptly clubbed at apex) and elongate setae on the genae, similar to those in the lyttine genus Lyttonyx . It diVers from the SW African Paractenodia (see below), which also has seven antennal segments, by the shape (subequal length and width dimensions) and position of the aedeagal hooks (distant from apex), the unmodi ed mesosternum and the absence of a distinct mesepisternal carina. After examining a paratype of this species, we are transferring it to Actenodia (new combination). This is primarily based on the structure of the antennae, male genitalia and mesosternum. Unlike most Actenodia , however, the pronotum is not strongly transverse and the mesepisterna are slightly furrowed although less so than in Paractenodia .

Eight S Mediterranean and Saharan species of this genus having nine antennal segments were historically assigned to Coryna , a synonym of Hycleus (see Bologna, 1991a: 275).

The rst-instar larvae of two S Mediterranean species have been described.