Fraudiziba mathiasi ( Bałuk, 1997 ) Harzhauser & Landau, 2021

Harzhauser, Mathias & Landau, Bernard, 2021, The Mitridae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea, Zootaxa 4983 (3), pp. 1-72 : 34-35

publication ID	https://doi.org/ 10.11646/zootaxa.4983.1.1
publication LSID	lsid:zoobank.org:pub:6A4778D6-195A-4AB1-AA1E-7D8000185B28
DOI	https://doi.org/10.5281/zenodo.5044084
persistent identifier	https://treatment.plazi.org/id/A82A87E9-8A37-386F-FF4D-F9A4FC24F97C
treatment provided by	Plazi
scientific name	Fraudiziba mathiasi ( Bałuk, 1997 )
status	comb. nov.

Treatment

Fraudiziba mathiasi ( Bałuk, 1997) View in CoL nov. comb.

Figs 9A View FIGURE 9 1 –A View FIGURE 1 2 View FIGURE 2 , B 1 –B View FIGURE 1 2 View FIGURE 2 , C 1 –C View FIGURE 1 2 View FIGURE 2 ; 10A View FIGURE 10 1 –A View FIGURE 1 2 View FIGURE 2

Mitra goniophora Bell. View in CoL — Hoernes & Auinger 1880: 77, pl. 7, fig. 9 [non Bellardi, 1850].

Mitra goniophora Var. View in CoL a— Hoernes & Auinger 1880: 78 (pars), pl. 9, fig. 12.

* Mitra goniophora Var. View in CoL a— Hoernes & Auinger 1880: 78, pl. 9, fig. 13.

Mitra (Mitraria) friedbergi hoernesi Mayer View in CoL — Kókay 1966: 62, pl. 9, fig. 7 [non Mayer, 1864].

Mitraria (Mitraria) mathiasi nom. n. — Bałuk 1997: 32 (pars) [non pl. 8, fig. 6].

Type material. Holotype: NHMW 2020 View Materials /0101/0001, SL: 31.3 mm, MD: 11.6 mm, Lăpugiu de Sus ( Romania), illustrated in Hoernes & Auinger (1880, pl. 9, figs 13a–b), figs 9B 1 –B 2.

Additional material. NHMW 1854/0035/0085, SL: 32.5 mm, MD: 10.5 mm, illustrated in Hoernes & Auinger (1880, pl. 7, figs 9a–b), figs 9A 1 –A 2, 10A 1 –A 2; NHMW 1854 View Materials /0035/0100, SL: 29.4 mm , MD: 10.5 mm, illustrated in Hoernes & Auinger (1880, pl. 9, figs 12a–b), figs 9C 1 –C 2; NHMW 2020 View Materials /0101/0003, SL: 33.4 mm , MD: 10.9 mm; NHMW 2020 View Materials /0101/0005, SL: 22.8 mm , MD: 8.1 mm; NHMW 2020 View Materials /0101/0006, 17 specimens, all Lăpugiu de Sus ( Romania) .

Revised description. Shell medium sized, moderately broad to moderately slender with narrow suture. Protoconch and first teleoconch whorl unknown. Teleoconch of at least nine whorls. Early teleoconch whorls weakly convex with periphery just above abapical suture, fifth to seventh whorls subcylindrical, with rounded or rarely angulated shoulder, coinciding with onset of spiral sculpture. Spiral sculpture of about 12 delicate, weakly raised spiral cords, separated by narrower interspaces. Spiral cords slightly more prominent below adapical suture and shoulder, nearly obsolete below shoulder. Last whorl subcylindrical, moderately high, slowly contracting. Spiral sculpture on shoulder persisting on last whorl, subobsolete on periphery. Base and fasciole with narrow primary and secondary spiral cords. Aperture moderately narrow to moderately wide, elongate, with narrowly incised anal canal. Columellar callus narrow, extending from adapical columellar fold to tip of siphonal canal. Columella with five oblique columellar folds, decreasing rapidly in strength abapically; fifth fold very weak or subosolete in most specimens. Outer lip thin. Siphonal canal moderately short, wide, slightly bent to the left with wide, moderately deep siphonal notch.

Colour pattern consisting of large subquadrate dots on shoulder and numerous thin spiral rows of black dashes below shoulder. Length of dashes alternating between spiral rows but constant within single spiral rows.

Shell measurements and ratios. SL: 20.3–33.4 mm, MD: 7.3–10.9 mm, AA: 39–41°, SL/ MD: 2.9–3.2, AL/ AW: 4.8–5.1, AH/S: 2.5–2.8.

Discussion. This species is quite variable concerning shape, ranging from moderately slender to stout shells. Bałuk (1997) referred to the stout specimen illustrated in Hoernes & Auinger (1880, pl. 9, figs 13a–b), which he designated as holotype. This designation was unfortunate as this specimen is an extreme morphotype. At first sight, the slender shell illustrated in Hoernes & Auinger (1880, pl. 7, fig. 9) seems to be too different to be conspecific with the broad holotype of F. mathiasi . Additional material from Lăpugiu de Sus ( Romania), however, reveals several intermediate specimens. Moreover, all specimens have the same delicate spiral sculpture and display identical colour pattern. The specimen from Lăpugiu de Sus illustrated in Hoernes & Auinger (1880, pl. 9, fig. 12) was treated by Davoli (2000) and Landau et al. (2013) as ‘ Ziba goniophora ’. Although, it agrees in outline with Fraudiziba goniophora , its sculpture is much more delicate and it lacks the punctate spiral grooves of F. goniophora . The morphology of its early teleoconch whorls and the sculpture, however, are identical with typical Z. mathiasi and therefore, we consider this specimen an angulated morphotype of Z. mathiasi .

The specimen from Korytnica ( Poland) illustrated by Bałuk (1997, pl. 8, fig. 6), as ‘ Mitraria mathiasi ’ is most probably unrelated to F. mathiasi . The Polish species differs in its angulated shoulder and, more importantly, lacks any spiral sculpture.

The colour pattern allows a clear separation from the Turkish, middle Miocene Fraudiziba ottomanica nov. sp., which has spiral rows of subquadrate dots. Fraudiziba paratethyca nov. nom. has fewer spiral rows of dots, lacks dashes and shows large subquadrate blotches on the shoulder.

Davoli (2000: 194) discussed a relationship of this species with Cancilla suballigata (Bellardi, 1887) but M. suballigata differs clearly in its conical spire and the prominent spiral sculpture (see syntype in Ferrero-Mortara et al. 1981, pl. 48, figs 12a–b).

Palaeoenvironment. Unknown. Probably middle to outer neritic environments.

Distribution in Central Paratethys. Badenian (middle Miocene): Pannonian Basin: Herend ( Hungary) ( Kókay 1966); Făget Basin: Lăpugiu de Sus ( Romania) ( Hoernes & Auinger 1880).