Anticyphon davidsoni, Ruta, Rafał, 2016

Ruta, Rafał, 2016, Anticyphon gen. nov., a new genus of Scirtidae (Coleoptera: Scirtoidea) inhabiting high altitude Andean cloud forests and páramo formation, Zootaxa 4175 (4), pp. 301-318 : 307-308

Anticyphon davidsoni sp. nov.

( Figs 1 A, 2A–C, 3A, 6)

Type material. Holotype, male ( CMNH): “ ECUADOR Loja . Cor- \ dillera Cordoncillo \ 11 km S Saraguro \ 3130 m. 27 Oct 1987 ”; “ C. Young, R. Davidson \ J. Rawlins. \ Cloud forest .” Paratype, male ( CMNH): “ ECUADOR: Chimborazo . \ 21 km ESE Licto, stream \ above Rio Alao. 3500m \ 18–19 October 1987 ”; “ J. Rawlins, C. Young \ R. Davidson. \ Montane woodland.”

Diagnosis. The largest species of the genus, pronotal disc with normal, not granulate punctures. Resembling A. santanderensis sp. nov. in size and pronotal punctation. It can be identified on the basis of male genitalia: trigonium is almost as long as parameroids (distinctly shorter in A. santanderensis sp. nov.).

Description. Male. Body oval, moderately large, slightly flattened, clothed with brownish procumbent setae. Colouration of dorsum brown, antennae and legs dark brown (holotype) or brown (paratype). Head small, 1.75× wider than interocular space, covered with very subtle granulate punctures; eyes relatively big, protuberant; frons with two shallow depressions. Antennae as in genus description. Mandibles without denticles on inner margins. Pronotum small, transversely rectangular, sides slightly curved, widest at posterior angles, anterolateral angles broadly rounded, not produced; disc moderately convex. Punctation of pronotum subtly granulate on lateral portions, very subtle and sparse in central portion. Pronotum without distinct pits along basal margin. Each elytron with 3 well marked longitudinal carinae. Elytral punctation irregular, relatively sparse; punctures shallow, separated by ca. 1.0 diameter. Penis ( Fig. 6 B) large (L 2.03 mm, W 0.42 mm), parameroids long, curved in apical portion, pointed at apices; trigonium subtriangular, without lateral processes, almost as long as parameroids, pala slightly shorter than parameroids, narrow, with membranous dorsal processes of penis; tegmen ( Fig. 6 C) large (L 1.52 mm, W 0.59 mm), with narrow, subtriangular parameres, pointed at apices; sternite VIII ( Fig. 6 D) small (L 0.40 mm, W 0.61 mm), widely V-shaped, with sparse setae in apical portions; sternite IX ( Fig. 6 E) relatively small (L 0.50 mm, W 0.65 mm), consisting of two subtriangular hemisternites, lightly sclerotized, with setose apical portion; tergite VIII ( Fig. 6 F) (L 0.98 mm, W 0.92 mm) with transversely rectangular apical plate, apical portion darker, covered with microsetae, apical margin with row of dense, short setae intermixed with sparse, longer ones, basal portion of apical plate with transverse rod, apodemes shorter than apical portion; tergite IX ( Fig. 6 G) (L 0.85 mm, W 0.50 mm) distinctly narrower than tergite VIII, central portion lightly sclerotized.

Female. Unknown.

Measurements and ratios. Males (n = 2): TL 6.67–6.75 (6.71) mm, PL 1.0–1.05 (1.03) mm, PW 2.15–2.25 (2.20) mm, EL 5.43–6.05 (5.74) mm, EW 3.78–3.80 (3.79) mm; TL/EW 1.76–1.78 (1.77), PW/PL 2.14–2.15 (2.15), EL/EW 1.44–1.59 (1.51), EL/PL 5.17–6.05 (5.61).

Distribution. Known from two localities in the southern part of Ecuadorian Andes ( Fig. 14 ).

Etymology. Named after Robert L. Davidson, curator of the Coleoptera section of the CMNH.

Figures

FIGURE 1. Anticyphon gen. nov., habitus. A) A. davidsoni sp. nov., B) A. ecuadorensis sp. nov., C) A. oyonensis sp. nov., female, D) A. oyonensis sp. nov., male E) A. paramoensis sp. nov., light form, F) A. paramoensis sp. nov., dark form, G) A. peruvianus sp. nov., H) A. santanderensis sp. nov. Scale bars = 1.0 mm.

FIGURE 6. Anticyphon davidsoni sp. nov., male genitalia. A) male genitalia, B) penis, C) tegmen, D) sternite VIII, E) sternite IX, F) tergite VIII, G) tergite IX. Scale bar = 0.5 mm. Abbreviations: dpp—dorsal process of penis, pe—penis, pm—paramere, pmd—parameroid, s—sternite, t—tergite, tg—tegmen, trg—trigonium.

FIGURE 14. Distribution of Anticyphon gen. nov. Please note that precise locality of A. peruvianus sp. nov. is unknown (see note under the species description).