Danainae

Bland, Keith P., 2019, Name-bearing types of butterflies (Lepidoptera, Papilionoidea), in the National Museums of Scotland, Edinburgh, Zootaxa 4559 (1), pp. 57-89 : 70-73

Danainae

Idea AGELIA Godart, in Latreille & Godart, 1819

Encyclopédie Méthodique Histoire Naturelle [Zoologie] 9. Entomologie i–328 [195]. Junior subjective synonym of Idea idea (Linnaeus, 1763) (see Grimshaw, 1897).

Syntypes, two males, pinned on strip of cork (one with left antenna and abdomen missing): Specimen data: Specimen reg.no.1; “Dufresne” [Printed] on verso “ Idea | agelia | 1” [Handwritten]. “ TYPE ” [Handwritten on oval pink label]. “Dufresne | 1936-50 | 1” on verso “Hestia idea Clerck | TYPE OF | Idea agelia Godart” [Handwritten (by Grimshaw) on mauve paper]. Specimen reg. no.2; Label as for no.1, only registration no.2. Remarks : Although not bearing Godart’s handwritten labels, these two specimens are unambiguously the two specimens in the Dufresne collection as seen by Godart and hence are syntypes. Grimshaw (1897) came to the same conclusion. The type locality is given as “les îles de Java & d’Amboine” [Islands of Java and Ambon (Moluccas)] .

Danais ALCATHOE Godart, in Latreille & Godart, 1819

Encyclopédie Méthodique Histoire Naturelle [Zoologie] 9. Entomologie i–328 [178]. Current accepted combination: Euploea alcathoe alcathoe (Godart, in Latreille & Godart, 1819) (see Ackery & Vane-Wright, 1984). Syntype, female, pinned underside uppermost (right antenna missing): Specimen data: “alcathoé fem. | alcathoe. encyc | amboine” [Handwritten (by Godart) on coarse bluish paper]. “Dufresne” [Printed] on verso “D.| alcathöe | fem. | 3” [Handwritten]. “TYPE” [Handwritten on oval pink label]. “Dufresne | 1936-50 | 7” on verso “ Euploea alcathoe | Godart | CO-TYPE?” [Handwritten (by Grimshaw) on mauve paper]. Remarks: This specimen, endorsed in Godart’s own hand, is unambiguously part of the type series. Grimshaw’s (1897) use of the term”probably” stems from his failing to recognise the significance of the original handwritten label. Type locality is “l’île d’Amboine” [Island of Ambon ( Moluccas)].

Danais ALOPIA Godart, in Latreille & Godart, 1819

Encyclopédie Méthodique Histoire Naturelle [Zoologie] 9. Entomologie i–328 [177–178]. Junior subjective synonym of Euploea midamus midamus (Linnaeus, 1758) (see Ackery & Vane-Wright, 1984).

Syntype, male, pinned underside uppermost (abdomen missing): Specimen data: “Dufresne” [Printed] on verso “D.| alopia | mas. | 7” [Handwritten]. “ TYPE ” [Handwritten on oval pink label]. “Dufresne | 1936-50 | 12” on verso “ Euploea midamus L. | CO-TYPE OF | alopia Godart” [Handwritten (by Grimshaw) on mauve paper]. Remarks: This is the sole survivor of the two specimens in the Dufresne Collection as seen by Godart. Although not endorsed

in Godart’s own hand it is undoubtedly part of the type series. The hesitation shown by Grimshaw (1897) is unfounded. Type locality is given as “l’île de Amboine” [Island of Ambon ( Moluccas)] but this may be an error.

Danais BAUDINIANA Godart , in Latreille & Godart, 1819

Encyclopédie Méthodique Histoire Naturelle [Zoologie] 9. Entomologie i–328 [181]. Junior subjective synonym of Euploea euphon (Fabricius, 1798) (see Ackery & Vane-Wright, 1984).

Syntypes, two males, pinned (one with abdomen and left antenna missing (No.8) and other with only left antenna missing (No.9)) Specimen data: Specimen reg. no.8; “De Baudin | Baudiniana | timor” [Handwritten (by Godart) on coarse bluish paper]. “Dufresne” [Printed] on verso “D. | Baudiniana | 2” [Handwritten]. “TYPE” [Handwritten on oval pink label]. “Dufresne | 1936-50 | 8” on verso “ Euploea euphon F. | TYPE OF | baudiniana | Godart” [Handwritten (by Grimshaw) on mauve paper]: Specimen reg.no.9; Labels as for no.8 above only no handwritten Godart label and registration no. 1936.50 9. Remarks: These are the original two specimens from the Dufresne Collection as seen by Godart and are unequivocally part of the type series. As enquiries at the Muséum National d’Histoire Naturelle, Paris failed to reveal likely type-specimens of this species (Ackery & Vane-Wright, 1984) the present specimens are the sole surviving syntypes. Type locality is given as “l’île de Timor [Island of Timor] par feu le capitaine Baudin.” Grimshaw (1897) correctly considered this locality to be in error as the species is found in the Mascarene islands. Interestingly Danais euphon is described by Godart on the same page of the Encyclopédie Méthodique but his description is merely copied from that of Fabricius .

Argynnis BRIAREA Godart, in Latreille & Godart, 1819

Encyclopédie Méthodique Histoire Naturelle [Zoologie] 9. Entomologie i–328 [261]. Current accepted combination: Anetia briarea (Godart, in Latreille & Godart, 1819) (see Ackery & Vane-Wright, 1984; Lamas, 2004).

Syntypes, one male and one female, pinned (female abdomen and most of both antennae missing): Specimen data: Specimen reg. no.30; “Dufresne” [Printed] on verso “ Argynnis | briarea | 1” [Handwritten]. “ TYPE ” [Handwritten on oval pink label]. “Dufresne | 1936-50 | 30” on verso “Clothilda | briarea Godart | CO-TYPE” [Handwritten (by Grimshaw) on mauve paper]. “ Argynnis briarea | Godart | SYNTYPE [Handwritten (by Lamas)] G.Lamas. det.1973” [Printed]: Specimen reg.no.31; “Briaréa. encyc.” [Handwritten (by Godart) on coarse bluish paper]. Other labels same as No.30 above only registration no. 1936-50.31. Remarks: According to the catalogue of the Dufresne collection, three specimens were present when the collection was studied by Godart. One of these specimens cannot now be traced. As concluded by Grimshaw (1897), the above two specimens are the remainder and so are part of the type series. Type locality is given as “Saint-Domingue” [Santo Domingo, Dominican Republic] .

Heliconia CLEOBAEA Godart, in Latreille & Godart, 1819

Encyclopédie Méthodique Histoire Naturelle [Zoologie] 9. Entomologie i–328 [222–223]. Current accepted combination: Lycorea halia (Hübner, 1816) ssp. cleobaea (Godart, in Latreille & Godart, 1819) (see Lamas, 2004). Syntypes, 1 male, pinned (left antenna missing) and 1 female mounted on cork strip and abdomen missing. Specimen data: Male “Dufresne” [Printed] on verso “ Heliconia | cleobaea | mas. 9” [Hamdwritten by Grimshaw]. “TYPE” [Oval pink label]. “Dufresne | 1936-50 | 14” [Handwritten by Grimshaw] on verso “ Lycorea | cleobaea Godart ♂ |?CO-TYPE” [Handwritten by Grimshaw]. “ Heliconia cleobaea | Godart | LECTOTYPE | G.Lamas det. 1973.” [Handwritten by G.Lamas on white card]: Female “Ceres C.”[Handwritten on bluish coarse paper]. “Cléobé fem.” [Handwritten by Godart on coarse paper]. “Dufresne” [Printed] on verso “ Heliconia | cleobaea | fem. 10” [Handwritten by Grimshaw]. “TYPE” [Oval pink label]. “Dufresne | 1936-50 | 15” on verso “ Lycorea | cleobaea Godart ♀ |?CO-TYPE” [All handwritten by Grimshaw]. “ Heliconia cleobaea | Godart | PARALECTOTYPE | G.Lamas det. 1973.” [Handwritten by Lamas].

Remarks: The lectotype and paralectotype designations have not been published (G. Lamas, pers.comm.)

Danais CLEOPHILE Godart , in Latreille & Godart, 1819

Encyclopédie Méthodique Histoire Naturelle [Zoologie] 9. Entomologie i–328 [185]. Current accepted combination: Danaus cleophile (Godart, in Latreille & Godart, 1819) (see Ackery & Vane-Wright, 1984; Lamas, 2004).

Syntype, male, pinned (abdomen missing): Specimen data: “Cléobula | Cleobula. encyc.” [Handwritten (by Godart) on coarse bluish paper]. “Dufresne” [Printed] on verso “D. | plexippus | 15” [Handwritten] - wrong label, according to Dufresne catalogue this is specimen no.13! “TYPE?” [Handwritten on oval pink label]. “Dufresne | 1936-50 | 5” on verso “Danaida cleophile | Godart | TYPE” [Handwritten (by Grimshaw) on mauve paper]. Remarks: Cléobule is listed, without any description, in the Section contents of the Encyclopédie Méthodique 9 ( Godart, 1819) as species No.32 of Danais . Species No.32 (p.185) is in fact called Danais cleophile suggesting that Godart had a late change of mind about the name of this species, but failed to revise the name in the Section contents and on the specimen data label. The above specimen labelled “Cléobula Cleobula. encyc.” by Godart is thus a syntype of Danais cleophile Godart, 1819 . No type locality is given, nor is there any indication how many specimens are involved in the type series. In spite of labelling the above specimen as a Type it is omitted from Grimshaw (1897). There is a female syntype in MNHN; the species is known from Hispaniola ( Haiti and Dominican Republic) and Jamaica (G. Lamas, pers.comm.).

Danais CLEOTHERA Godart, in Latreille & Godart, 1819

Encyclopédie Méthodique Histoire Naturelle [Zoologie] 9. Entomologie i–328 [185]. Current accepted combination: Danaus gilippus (Cramer, 1775) ssp. cleothera (Godart, in Latreille & Godart, 1819) (see Lamas, 2004).

Syntype, female, pinned: Specimen data: “Cleothère | Cleothera. encyc.” [Handwritten (by Godart) on coarse bluish paper]. “Dufresne” [Printed] on verso “D. | cleothera | 14” [Handwritten]. “ TYPE ” [Handwritten on oval pink label]. “Dufresne | 1936-50 | 6” on verso “Danaida | cleothera Godart | TYPE ” [Handwritten (by Grimshaw) on mauve paper]. Remarks: This specimen is unambiguously the single specimen in the Dufresne collection as seen by Godart and is thus a syntype. The type locality is given as “l’île de Timor” [ Island of Timor ] but according to Grimshaw (1897) it is erroneous as this subspecies is known from Hispaniola ( Haiti and Dominican Republic) and Puerto Rico .

Heliconia EUCLEA Godart , in Latreille & Godart, 1819

Encyclopédie Méthodique Histoire Naturelle [Zoologie] 9. Entomologie i–328 [220–221]. Current accepted combination: Hypothyris euclea euclea (Godart, in Latreille & Godart, 1819) (see Lamas, 2004).

Holotype, female, pinned on strip of cork (antennae missing). Specimen data: “Eucléa, encyc” [Handwritten (by Godart) on coarse creamy paper]. “Dufresne” [Printed] on verso “Heloconia | euclea | 12” [Handwritten]. “TYPE” [Handwritten on oval pink label]. “Dufresne | 1936-50 | 17” on verso “Ceratinia | euclea Godart | TYPE” [Handwritten (by Grimshaw) on mauve paper]. “ Heliconia euclea | Godart | HOLOTYPE | [Handwritten (by Lamas)] G.Lamas. det. 1973 [Printed]”. Remarks: As indicated by Grimshaw (1897) this specimen is unambiguously part of the type series of Heliconia euclea Godart, 1819 . Type locality is given as “Antilles” [The Antilles, West Indies]. As Lamas consulted the Paris Museum collection before visiting Edinburgh in 1973 he is probably correct in implying that this specimen is the only one extant.

Euploea LIMBORGII Moore, 1878 [1879]

Proceedings of the Zoological Society of London for 1878 (4), 821–859 [823], plate LI, figure 2. Current accepted combination: Euploea algea (Godart, in Latreille & Godart, 1819) ssp. limborgii Moore,1878 [1879] (see Morishita, 1970).

Syntypes, 2 males, pinned. Specimen data: Specimen reg. no.10; “Ahsown”[Handwritten]. “ Euploea Limborgii ♂ | (Type) Moore”[Handwritten]. “Tenasserim | 1936-50 | 10”on verso “ Euploea limborgii | Moore CO-TYPE”[Handwritten on mauve paper]. Specimen reg. no.11; “Hatsiega, Homydaron”[Handwritten]. “ Euploea Limborgii ♂ | (Type) Moore”[Handwritten]. “Tenasserim | 1936-50 | 11” on verso “ Euploea limborgii | Moore CO-TYPE” [Handwritten on mauve paper]. Remarks : These specimens are obviously part of the type series collected on the Upper Tenasserim Expedition of 1876-1877. The species is illustrated in colour in figure 2 of plate 51 ( Moore, 1878 [1879]) .

Danais MEGANIRE Godart, in Latreille & Godart, 1819

Encyclopédie Méthodique Histoire Naturelle [Zoologie] 9. Entomologie i–328 [192]. Current accepted combination: Ideopsis juventa (Cramer, 1777) ssp. meganire (Godart, in Latreille & Godart, 1819) (see Tsukada,

Yata & Morishita, 1985). However, Vane- Wright (1975) regards meganire as a junior subjective synonym of subspecies claviger (Gmelin, [1790]).

Paralectotypes, 2 males, pinned, both designated by R.I.Vane-Wright, 1974 ( Vane-Wright, 1975). The lectotype is in the Muséum National d’Histoire Naturelle ( MNHN) in Paris .

Heliconia MEGARA Godart, in Latreille & Godart, 1819

Encyclopédie Méthodique Histoire Naturelle [Zoologie] 9. Entomologie i–328 [223]. Current accepted combination: Tithorea harmonia (Cramer, 1777) ssp. megara (Godart, in Latreille & Godart, 1819) (see D’Abrera, 1984; Lamas, 2004).

Syntype, male, pinned (repinned on to strip of cork and head missing): Specimen data: “mégare | megara”. [Handwritten (by Godart) on coarse creamy paper]. “Dufresne” [Printed] on verso “ Heliconia | megara | 6” [Handwritten]. “Dufresne | 1936-50 | 16” on verso “Hirsutis harmonia | Cram. | form megara Godart |?CO-TYPE” [Handwritten (by Grimshaw) on mauve paper]. “ Heliconia megara | Godart | LECTOTYPE [Handwritten by Lamas] | G.Lamas. det. 1973 [Printed]”. Remarks: This is the specimen present in the Dufresne collection when it was studied by Godart and the specimen is named in Godart’s handwriting. It is thus part of the type series. The lectotype designation has not been published (G. Lamas pers.comm.).Type locality is given as “les Antilles” [The Antilles].

Danais PROTHOE Godart, in Latreille & Godart, 1819

Encyclopédie Méthodique Histoire Naturelle [Zoologie] 9. Entomologie i–328 [177]. Junior subjective synonym of Euploea phaenareta phaenareta (Schaller, 1785) (see Vane-Wright, 1975).

Lectotype, male, pinned on cork strip (both antennae missing): Specimen data: “prothoé. mâle | midamus. Cram | amboine” [Handwritten (by Godart) on coarse bluish paper]. “Dufresne” [Printed] on verso “D. | prothöe | mas, | 6” [Handwritten]. “Dufresne | 1936-50 | 13” on verso “ Euploea phaenarete | Schaller | TYPE OF | prothoe Godart” [Handwritten (by Grimshaw) on mauve paper]. “LECTO | TYPE” [round, red-bordered label]. “ Danais prothoe | Godart | LECTOTYPE ♂ [Handwritten (by Vane-Wright)] | det. R.I.Vane-Wright 1974 [Printed]”. Remarks: This specimen is the sole survivor of the 2 specimens from the Dufresne collection seen by Godart and is thus part of the type series. Grimshaw (1897) came to the same conclusion. It was designated as the lectotype in Vane-Wright (1975) and labelled accordingly. Type locality is “l’île d’ Amboine” [Isle of Ambon ( Moluccas)].