Pristapanesiinae Engel et al., 2013

Lepeco, Anderson & Melo, Gabriel A. R., 2024, Revisiting the phylogeny of the scolebythid wasps (Hymenoptera: Aculeata) through Bayesian model evaluation and parsimony, with description of a new fossil family of Chrysidoidea, Zoological Journal of the Linnean Society 201, pp. 57-85 : 71

Subfamily Pristapanesiinae Engel et al., 2013

Type genus: Pristapenesia Brues, 1933 .

Included genera: † Acanthabythus Lepeco and Melo, 2022a ; † Arcapenesia Lepeco and Melo, 2022a ; † Burmapenesia Lepeco and Melo, 2022 ; † Cephalobythus Lepeco and Melo, 2022a ; † Eobythus Lacau et al., 2000 ; † Gnathapenesia Lepeco and Melo, 2022a ; Pristapenesia Brues, 1933 ; † Uliobythus Engel and Grimaldi, 2007 ; † Zapenesia Engel and Grimaldi, 2007 .

Diagnosis. Pedicel with two distinctly long setae, longer than apical width of pedicel, in addition to short setae; dorsal portion of eye distinctly narrowed in relation to ventral one; hypostomal bridge very long, more than 2.0× basal width of mandible; metapostnotum not visible externally or with medial portion extending onto propodeum; propodeal spiracle contiguous with metapleural line; longest setae on metatibia longer than 2× maximum tibial width; forewing with submarginal and medial cells not enclosed by tubular veins; tubular portion of vein C restricted to base of hindwing.

Taxa presently removed from Scolebythidae

In recent years, many fossil taxa have been assigned to Scolebythidae , but some of them belong to other aculeate groups and are here formally removed from Scolebythidae . Except for † Boreobythus , we have not examined their respective type material and our interpretation is based mainly on the descriptions and images presented in the original articles and on the results of the present analyses (see Supporting Information, File C, Figs S1 , S 2 ). In the case of taxa described from Burmese amber, we also had the opportunity to study additional inclusions that fit the described taxa. In summary, the genera † Boreobythus and † Cursoribythus were moved to Bethylidae , while the genera † Ectenobythus , † Necrobythus , and † Sphakelobythus were moved to Chrysididae . Characters linking those taxa to their respective families are listed in Appendix 2.

Figures

Figure 1. Phylogenetic tree showing the relationships obtained from parsimony analysis under implied-weighting (k = 3). Black and white hashmarks indicate unique and homoplastic changes, respectively.Only unambiguous changes are shown. Habitus of †Chrysopsenella euryphaessa gen.et sp. nov.shown within blue frame.Habitus of †Cephalobythus clypeatus Lepeco and Melo, 2022 shown within orange frame.

Figure 2. Phylogenetic trees obtained from Parsimony analyses.A, under equal weights. B, under implied weighting (k = 3).