Angustopila apiaria Pall-Gergely & Hunyadi, 2023

Pall-Gergely, Barna, Hunyadi, Andras, Vermeulen, Jaap J., Grego, Jozef, Sutcharit, Chirasak, Reischuetz, Alexander, Dumrongrojwattana, Pongrat, Botta-Dukat, Zoltan, Oerstan, Aydin, Fekete, Judit & Jochum, Adrienne, 2023, Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae), ZooKeys 1147, pp. 1-177 : 1

Angustopila apiaria Pall-Gergely & Hunyadi sp. nov.

Fig. 86

Type material.

Holotype: Vietnam • 1 empty shell (H: 1.04 mm, D: 0.85 mm); Ðà Nẵng, Ngũ Hành Sơn, Thái Sơn, vicinity of Chùa Quán Thế Äm (locality code: 2019/25a); 15°59.94'N, 108°15.33'E; 10 m a.s.l.; 11 Feb. 2019; A. Hunyadi leg.; HNHM 105316.

Paratypes: Vietnam • 37 shells; same data as for holotype; coll. HA • 3 shells; same data as for holotype; coll. JJV.

Additional material.

Vietnam • 4 j/b shells; same data as for holotype; coll. HA.

Diagnosis.

A medium-sized, high conical Angustopila species with a narrow umbilicus, protruding aperture, widely spaced and strong spiral striation, and three prominent apertural barriers (slightly curved parietal, deeply situated lower palatal, inward running subcolumellar).

Description.

Shell of normal size for the genus, higher than wide; off-white, conical with slowly increasing whorls; body whorl widest from standard apertural view; protoconch consists of 1.25 whorls, with very slight indication of spiral striation preceding the first teleoconch whorl; teleoconch ornamented by fine, irregular and weak, radial growth lines and conspicuous, prominently raised and equidistantly-spaced spiral striae (ca. 12 or 13 on body whorl from standard apertural view); whorls 4.75, rounded; aperture only slightly oblique to shell axis from lateral view; umbilicus moderately wide; aperture protruding obliquely (visible from basal view), aperture subquadrate with straight parietal side; peristome expanded, not reflected; parietal callus protruding, detached from penultimate whorl; parietal tooth tongue-like and extended, its base straight, its tip slightly turned up in palatal direction, but bent away from palatal tooth; lower palatal tooth low but prominent, wide, elongated, almost perpendicular to the peristome, its outer end situated close to basal lip; subcolumellar tooth strong, as high as the palatal but less wide, elongated, perpendicular to peristome, its outer end situated on the lower part of columellar side of peristome.

Measurements (in mm).

H = 0.93-1.04; D = 0.8-0.86, H/D*100 = 113.4-126.3 (n = 6), RUD = 25.9-27.2 (n = 2).

Differential diagnosis.

The high conical shell shape, narrow umbilicus, protruding aperture, prominent, widely spaced spiral striae, and three strong apertural teeth distinguish this species from all other Angustopila species. The most similar species is Angustopila tridentata sp. nov., which is concave-conical instead of ovoid or high-conical and bears a wider umbilicus and denser spiral striation. The columellar tooth of A. apiaria sp. nov. runs perpendicular to the shell axis (runs inside the aperture) whereas that of A. tridentata sp. nov. runs along the peristome. The parietal tooth of A. apiaria sp. nov. is located deeper and positioned lower than that of A. tridentata sp. nov.

Etymology.

Named after its shape reminiscent of a classic apiary, beehive (apiarium) in Latin.

Distribution.

This species is known only from the type locality region of Chùa Quán Thế Äm, Vietnam (Fig. 19 ).

Remarks.

This is the only Angustopila species possessing teeth other than the parietal lamella (lower palatal and subcolumellar teeth) that are perpendicular to the peristome, i.e., run into the aperture. This may mean that A. apiaria sp. nov. would deserve a genus of its own. Moreover, this is the most south-eastern occurring Angustopila species. Central Vietnam so far seems to have different land snail fauna than northern Vietnam.

Figures

Figure 19. Distribution of Angustopila species.

Figure 86. Angustopila apiaria Pall-Gergely & Hunyadi, sp. nov. A paratype (specimen 2) B-I holotype (HNHM 105316). Apertural (A, B), lateral (C), ventral (D), and apical (E) sides of the shell; aperture (F); sculpture on the protoconch (G), ventral (H) and frontal (I) surface of the body whorl.