Harperocallis McDaniel, J. Arnold Arbor. 49 (1): 36. 1968.

Campbell, Lisa M. & Dorr, Laurence J., 2013, A synopsis of Harperocallis (Tofieldiaceae, Alismatales) with ten new combinations, PhytoKeys 21, pp. 37-52 : 37

Harperocallis McDaniel, J. Arnold Arbor. 49 (1): 36. 1968. Figs 1 -3

Type.

Harperocallis flava McDaniel.

Key to the Species of Harperocallis

(modified from: Cruden 1991, and Campbell 2010)

1	Capsules ribbed or unribbed, well-developed lateral veins never extending to the style base	6
-	Capsules 9-ribbed, well-developed lateral veins extending to the style base	2
2	Inflorescence compound, the branches subtended by chlorophyllous cataphylls	Harperocallis paniculata
-	Inflorescence a simple raceme; peduncular bracts scarious	3
3	Flowers pendant	Harperocallis penduliflora
-	Flowers erect	4
4	Leaves 2.0-4.0 mm wide, usually glabrous, occasionally ciliate at the sheath apex; peduncles <1.5 mm wide; calycular bracts longer than wide; tepals 6.0-9.0 × 1.5-2.0 mm; anthers 0.9-1.1 mm long	Harperocallis duidae
-	Leaves 3.0-7.0 mm wide, margin ciliolate to hirsutulous; peduncles> 1.5 mm wide; calycular bracts usually wider than long, sometimes equal; tepals 11-14 × 2-4 mm; anthers> 1.1 mm long	5
5	Leaves 3.0-6.5 mm wide; peduncular bracts 3-8 (-10); inflorescence ca. 9-30-flowered; tepals enclosing the rest of the flower; anthers 1.2-1.8 mm long	Harperocallis schomburgkiana
-	Leaves 6.0-7.0 mm wide; peduncular bracts 3 or 4; inflorescence ca. 25-40-flowered; tepals open; anthers 2.0-2.5 mm long	Harperocallis neblinae
6	Capsule incompletely 9-ribbed, well-developed lateral veins never extending to the style base	7
-	Capsule 3-ribbed or unribbed	8
7	Inflorescence with eglandular hairs; peduncular bracts (3-) 4-11 (-15); flowers (4-) 8-28; styles recurved, apices oblique; stigmas facing outward; capsule etuberculate	Harperocallis sessiliflora
-	Inflorescence glabrous; peduncular bracts 7-17; flowers 23-35; styles straight; stigmas capitate; capsule tuberculate	Harperocallis robustior
8	Leaves glabrous, veins not prominent; inflorescence 1-flowered; flower erect; ovary densely tuberculate; capsule unribbed	Harperocallis flava
-	Leaf margin tomentose, ciliolate or glabrous, veins prominent; inflorescence> 1-flowered, or if 1-flowered, the flower pendant; ovary not tuberculate; capsule 3-ribbed	9
9	Peduncular bract solitary, chlorophyllous; inflorescence 1-4-flowered; flowers pendant	Harperocallis sipapoensis
-	Peduncular bracts (1) 2-8, scarious; inflorescence> 10-flowered; flowers erect	10
10	Calycular bracts usually longer than wide (1.8-3.0 × 1.7-2.5 mm); styles 0.3-0.8 (-0.9) mm long, straight; stigmas capitate	Harperocallis falcata
-	Calycular bracts usually wider than long (1.5-2 × 1.8-2.3 mm); styles 0.9-1.1 mm long, recurved, apices oblique; stigmas facing outward	Harperocallis longiflora

Figures

Figure 1. Distribution of Harperocallis.

Figure 3. Harperocallis robustior. A Habit B Leaf (detail of parallel veins and tomentose margins) C Inflorescence (detail showing peduncular and calycular bracts subtending flowers) D Peduncular bracts (detail) E Flower F Tepal (adaxial view) G Ovary (note the tuberculae) H Ovary (cross-section showing axile placentation and ovules) I Capsule (carpels separating distally). (A, I from L. J. Dorr et al. 4967 (US); B-H from B. Stergios et al. 17327 (PORT).