Incoltorrida benesculpta, Perkins & Bergsten, 2019

Perkins, Philip D. & Bergsten, Johannes, 2019, New Myxophagan water beetles from Madagascar (Coleoptera: Torridincolidae, Hydroscaphidae), Zootaxa 4657 (1), pp. 57-96: 72-73

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Incoltorrida benesculpta

new species

Incoltorrida benesculpta   , new species

Figs. 8 View FIGURE 8 (habitus); 16, 20 (genitalia); 35 (map); 42, 43 (habitat)

Type Material. Holotype (male): Fianarantsoa, 3.2km S Ambohimanjaka , 20º 14.0343’ S 47º 5.59145’ E, waterfall and hygropetric habitat near hwy. 7, elev. 1415 m, 5 xi 2014, P. D. Perkins ( NHRS) GoogleMaps   . Paratypes (19): Same data as holotype (13: NHRS, MCZ, BMNH, PBZT / MBC); Antsiranana, Sava, Mad.; Antsirana ; Marojejy NP, 800m N from Camp I, Humbert waterfall; hygropetric; 14.4333S 49.773E; MAD 14-48 View Materials GoogleMaps   , ex. bedrock pools at side of waterfall, elev. 550 m, 8 xi 2014, P. D. Perkins ( MAD 14-48 View Materials ) (6 MCZ)   .

Differential Diagnosis. Together with I. galoko   the smallest known member of the genus (l/w ca. 1.85/1.05). The elytral costae are high and strong, and are more sinuate than in other species. The serial punctures are large and in sulcate grooves. The sculpture of the pronotum is similar to that of I. quintacostata   , but I. benesculpta   has a more distinct midlongitudinal carina in the basal half, and the sides of the pronotum in the anterior half are more arcuate. The elytra and the aedeagi of the two species are quite dissimilar.

Description. Size: holotype (length/width, mm): body (length to elytral apices) 1.85/1.05; head width 0.40; pronotum 0.40/0.81; elytra 1.17/1.05. Dorsum black, venter dark brown, legs brown except femoral-tibial articulations and tarsi black.

Sides of frontoclypeal shield slightly arcuate toward the midline, such that apical part is slightly wider than the width at midlength, and slightly narrower than the posterior area of the frons.

Short oblique carinae on anterior 1/3 of pronotum are well developed, no indication of midlongitudinal carina. The basal 1/3 has a short carina on each side, and a very low narrow midlongitudinal carina.

Elytron with eight strong high costae; fourth costa interrupted with strong punctures; fifth costa weakly bisinuate; sixth and seventh costae very indistinct. Serial punctures large and distinct.

Sides of metaventral tabella more distinctively arcuate than in other species. Midlongitudinal groove in the posterior 1/2 of metaventral tabella gradually widens from posterior to anterior.

Midlongitudinal carina of first abdominal ventrite strong and extends length of ventrite.

Male genitalia very short, with the apical part narrowed and spinose ( Figs. 16 View FIGURES 14–18 , 20 View FIGURES 19–20 ).

Etymology. Named in reference to the markedly sculptured pronotum and elytra.


Swedish Museum of Natural History, Entomology Collections


Museum of Comparative Zoology


Parc Botanique et Zoologique de Tsimbazaza


Montgomery Botanical Center


Madras Museum