APHODIINAE

Peck, Stewart B., 2011, The diversity and distributions of the beetles (Insecta: Coleoptera) of the northern Leeward Islands, Lesser Antilles (Anguilla, Antigua, Barbuda, Nevis, Saba, St. Barthélemy, St. Eustatius, St. Kitts, and St. Martin-St. Maarten, Insecta Mundi 2011 (159), pp. 1-54 : 22-23

SUBFAMILY APHODIINAE

TRIBE APHODIINI

Nialaphodius nigrita (Fabricius) 1801: 73 ( Aphodius ); Gordon and Skelley 2007: 265; Ivie et al. 2008b: 244 ( Aphodius ). = A. cuniculus Chevrolat 1864: 411 , Chapin 1940: 7; Ramos 1946: 41; Wolcott 1950: 248 of Puerto Rico, of Mona, of Vieques; synonymy in Bordat 1990: 62; Paulian 1947a: 37 ( Nialus ); Chalumeau and Gruner 1974: 795; Cartwright and Chalumeau 1978: 8; Bennett and Alam 1985: 22; Woodruff et al. 1998: 32; Chalumeau 1983a: 58 (all as Aphodius ( Nialus )); Valentine and Ivie 2005: 276; generic synonymy in Skelley et al. 2007; Turnbow and Thomas 2008; 49. = A. granarius variety guadeloupensis Fleutiaux and Sallé 1890: 395 . Distribution. Antigua, Barbados, Carriacou, Cuba, Désirade, Dominica, Grenada, Guadeloupe, Guana, Hispaniola, Les Saintes, Jamaica, Marie-Galante, Martinique, Mona, Montserrat, Mustique, Puerto Rico, St. Croix, St. John, St. Kitts, St. Lucia, St. Thomas, St. Vincent, Tortola, Vieques. Widespread southern USA, Mexico to widespread South America , Tobago; introduced to Lesser Antilles. Introduced to New World from Afro-tropical region; worldwide in tropics and subtropics. Notes. Common in cow dung.

TRIBE EUPARIINI

Ataenius gracilis (Melsheimer) 1845: 137 ( Oxyomus ); Fleutiaux and Sallé 1890: 397; Chapin 1940: 25; Paulian 1947a: 44; Chalumeau and Gruner 1974: 806; Cartwright and Chalumeau 1978: 12; Chalumeau 1978: 44, 1983a: 74; Bennett and Alam 1985: 22; Woodruff et al. 1998: 33; Stebnicka 2007b: 79; Ivie et al. 2008b: 244; Turnbow and Thomas 2008; 48. = A. chilensis Solier 1851: 72 of Barbados in Woodruff et al. 1998: 33. Distribution. Antigua, Bahamas, Barbados, Cuba, Dominica, Grenada, Guadeloupe, Hispaniola, Jamaica, Marie-Galante, Martinique, Montserrat, Puerto Rico, St. Croix, St. Kitts, St. Vincent, Vieques. United States, Canada; throughout the Neotropics (Argentina, Chile, Colombia, Peru, Galapagos Islands); widespread New World. Notes. Adults attracted to lights; found in cow dung.

Ataenius howdeni Chalumeau 1978: 51 ; Stebnicka 2002: 269, 2007b: 21; Ivie et al. 2008b: 244. Distribution. Antigua, Montserrat; Lesser Antilles endemic.

Ataenius liogaster Bates 1887: 94 ; Chapin 1940: 29; Paulian 1947a: 44; Chalumeau and Gruner 1974: 813; Stebnicka and Lago 2005: 60 (synonymy); Stebnicka 2007b: 36; Ivie et al. 2008b: 244. = A. edwardsi Chapin 1940: 26 ; Cartwright and Chalumeau 1978: 14; Chalumeau 1983a: 82, 174; Bennett and Alam 1985: 22; Woodruff et al. 1998: 33. Stebnicka and Lago 2005: 60. Distribution. Antigua, Barbados, Carriacou, Cuba, Dominica (overlooked in Stebnicka and Lago 2005: 61), Grenada, Guadeloupe, Hispaniola, Jamaica, Montserrat, Puerto Rico, St. Croix, St. Lucia, St. Vincent. Mexico to Panama, Ecuador (including Galapagos), Trinidad, Venezuela; Micronesia; widespread Antilles and Latin America . Notes. Adults attracted to lights; found in cow dung.

Ataenius luteomargo Chapin 1940: 36 ; Paulian 1947a: 41; Chalumeau and Gruner 1974: 811; Cartwright and Chalumeau 1978: 12; Chalumeau 1983a: 84; Bennett and Alam 1985: 22; Woodruff et al. 1998: 33; Stebnicka 2002: 270, 2007b: 22. = A. terminalis Fleutiaux and Sallé 1890: 396 ; Uyttenboogaart 1902: 116 of Barbados; Arrow 1903: 512 (not Chevrolat). = A. versicolor Hinton 1937: 183 (not Schmidt) of Barbados. Distribution. Antigua, Barbados, Désirade, Dominica (type locality), Grenada, Guadeloupe, Hispaniola, Les Saintes, Marie-Galante, Martinique, Montserrat, Puerto Rico, St. Kitts, St. Lucia; not Jamaica. Panama, Surinam, Venezuela; widespread Antilles and Latin America . Notes. Adults attracted to lights, found in cow dung.

Ataenius picinus Harold 1867: 281 ; Chalumeau and Gruner 1974: 810; Cartwright and Chalumeau 1978: 14; Chalumeau 1978: 44, 1983a: 80; Stebnicka 2004: 224, 2007b: 42. = A. elongatus Beauvois 1811: 104 , Fleutiaux and Sallé 1890: 397 of Guadeloupe. = A. darlingtoni Hinton 1937: 179 ; Chapin 1940: 30; Ramos 1946: 41; Paulian 1947a: 40. Distribution. Antigua, Cuba, Dominica, Grenada, Guadeloupe, Hispaniola, Jamaica, Marie-Galante, Martinique, Mona, Puerto Rico, St. Croix, St. Vincent. widespread New World; nearly cosmopolitan; United States, Argentina, Brazil, Uruguay; Australia, Fiji, New Caledonia, New Hebrides, New Zealand. The most widely distributed species in the genus, and undoubtedly on all islands of the Lesser Antilles. Notes. Adults attracted to lights; found in cow dung.

Ataenius scabrellus Schmidt 1909: 118 ; Chalumeau and Gruner 1974: 807; Stebnicka 2003: 227, 2007b: 27; Ivie et al. 2008b: 244; Turnbow and Thomas 2008; 48. = A. havanensis Balthasar 1938: 56 ; Chalumeau and Gruner 1974: 807; Chalumeau 1983a: 75. = A. miamii Chapin 1940: 41 ; Ramos 1946: 41; Bennett and Alam 1985: 22; Tucker 1952: 342 of Barbados; Miskimen and Bond 1970: 97 of St. Croix. = A. fleutiauxi Paulian 1947a: 42 of Guadeloupe. Distribution. Bahamas, Barbados, Cuba, Bahamas, Désirade, Hispaniola, Jamaica, Les Saintes, Puerto Rico, Guadeloupe, Martinique, Mona, Montserrat, St. Croix, St. Kitts. USA (FL); Venezuela, Guyana; widespread Antilles and Latin America .

Ataenius scutellaris Harold 1867: 82 ; Chalumeau and Gruner 1974: 803; Cartwright and Chalumeau 1978: 14; Chalumeau 1978: 44, 1983a: 69; Bennett and Alam 1985: 22; Valentine and Ivie 2005: 276; Stebnicka 2007a: 48, 2007b: 59; Ivie et al. 2008b: 244. = Ataenius frater Arrow 1903: 512 of St. Vincent; Chapin 1940: 32; Woodruff et al. 1998: 33. = A. elongatus Beauvois 1811: 104 ; Fleutiaux and Sallé 1890: 397 of Guadeloupe. Distribution. Antigua, Bahamas, Barbados, Cuba, Dominica, Grenada, Guadeloupe, Guana, Hispaniola, Jamaica, Marie-Galante, Martinique, Montserrat, Puerto Rico, St. Croix, St. Kitts, St. Lucia, St. Martin-St. Maarten, St. Thomas, St. Vincent, Tortola. Mexico to Bolivia and Brazil, Trinidad; widespread Antilles and Latin America ; introduced to Vanuatu (New Hebrides), Seychelles, Mascarenes, Malaya. Notes. Adults attracted to lights; found in cow dung.

Ataenius temperei Chalumeau and Gruner 1974: 799 ; Cartwright and Chalumeau 1978: 12; Chalumeau 1983a: 64; Stebnicka 2007b: 69; Ivie et al. 2008b: 244. Distribution. Dominica, Guadeloupe (type locality), Montserrat, St. Kitts; Lesser Antilles endemic. Notes. Adults attracted to lights; not found on cow dung.