Paraphloeostiba, Steel, 1960

Shavrin, Alexey V., 2020, New species and records of Paraphloeostiba Steel, 1960 from China and Laos and descriptions of four new species of related genera (Coleoptera: Staphylinidae Omaliinae: Omaliini), Zootaxa 4890 (3), pp. 301-329 : 317

Key to species of Paraphloeostiba of China and Laos

1 Pronotum slightly flattened, with acute posterior angles....................................................... 2

- Pronotum convex, with obtuse posterior angles.............................................................. 6

2 Parameres wide, significantly extending beyond apex of median lobe............................................ 3

- Parameres short, not reaching apex of median lobe........................................................... 5

3 Median lobe very wide ( Figs. 1–2 ). Body brown. Female accessory sclerite as in Fig. 10 . Body length: 2.00– 2.50 mm. China (Yunnan).................................................................................... P. ampliata

- Median lobe narrower. Body slightly darker................................................................ 4

4 Parameres very long, narrow in apical portions, with several moderately long apical setae (see figure 21 in Shavrin & Smetana 2016). Apical portion of female accessory sclerite narrow ( Shavrin & Smetana 2016: figure 11). Body length: 2.10–2.30 mm. Myanmar, Taiwan............................................................................... P. sonani

- Parameres shorter, wider in apical portion, with very long apical and preapical setae ( Fig. 4 ). Apical portion of female accessory sclerite moderately wide ( Steel 1960: figure 81). Body length: 1.90–2.10 mm. Indonesia (Sumatra), China ( Hong Kong).............................................................................................. P. sulcicollis

5 Pronotum narrow, without impression in medioapical third. Body brown. Apex of median lobe narrow, acute ( Figs. 15, 17 ). Female accessory sclerite with wide and short apical portion ( Fig. 22 ). Body smaller: 1.50–1.82. Laos........ P. brancuccii

- Pronotum disitnctly transverse, with narrow impression in medioapical third. Body yellow-brown to reddish-brown. Apex of median lobe moderately wide, rounded ( Figs. 28, 30 ). Female accessory sclerite with narrow and long apical portion ( Fig. 35 ). Body larger: 2.13–3.32. Vietnam, Laos.......................................................... P. opacicollis

6 Pronotum from middle more sharply narrowed posteriad than anteriad. Body brown. Parameres not reaching apex of median lobe ( Fig. 23 ). Body length: 1.90 mm. China (Zhejiang)................................................ P. cooteri

- Pronotum from middle evenly narrowed both anteriad and posteriad. Coloration different. Parameres extending beyond apex of median lobe......................................................................................... 7

7 Punctation of pronotum and elytra moderately fine and sparse. Head, pronotum, and abdomen brown, elytra yellow-brown. Median lobe slightly narrowed apically; parameres narrow ( Fig. 25 ). Female accessory sclerite with narrow and long apical portion ( Steel 1960: figure 99). Body length: 1.70–3.30 mm. Europe, China, North America, Australia, New Zealand.............................................................................................. P. gayndahensis

- Punctation of pronotum and elytra deep and dense. Body yellow-brown to red-brown. Median lobe significantly narrowed apically; parameres wide ( Fig. 5 ). Female accessory sclerite with wide and short apical portion ( Shavrin & Smetana 2016: figure 5; Shavrin 2017: figure 17). Body length: 1.80–2.50 mm. China, Taiwan, Vietnam........................ P. formosana

Figures

FIGURES 1–5. Aedeagus (parameral view) of Paraphloeostiba: 1–2—P. ampliata (1: paratype; 2: holotype), 3—P. sp. (Hong Kong), 4—P. sulcicollis (Hong Kong), 5—P. formosana (Vietnam). Scale bar: 0.1 mm.

FIGURES 6–13. Abdominal segments of Paraphloeostiba (6–11: P. ampliata; 12–13: P. cooteri): 6, 12—male tergite VIII (dorsal view), 7, 13—male sternite VIII (ventral view), 8—female tergite VIII (dorsal view), 9—female sternite VIII (ventral view), 10—female accessory sclerite, 11—spermatheca. Scale bar: 0.1 mm.

FIGURES 15–17. Aedeagus of Paraphloeostiba brancuccii (15, 17: parameral view, 16: lateral view): 15–16—paratype, Xieng Khouang, 17—paratype, Bolikhamsay. Scale bar: 0.1 mm.

FIGURES 18–22. Abdominal segments of Paraphloeostiba brancuccii: 18—male tergite VIII (dorsal view), 19—male sternite VIII (ventral view), 20—female tergite VIII (dorsal view), 21—female sternite VIII (ventral view), 22—female accessory sclerite. Scale bar: 0.1 mm.

FIGURES 28–30. Aedeagus of Paraphloeostiba opacicollis (Laos): 28, 30: parameral view, 29—lateral view. Scale bar: 0.1 mm.

FIGURES 31–35. Abdominal segments of Paraphloeostiba opacicollis: 31—male tergite VIII (dorsal view), 32—male sternite VIII (ventral view), 33—female tergite VIII (dorsal view), 34—female sternite VIII (ventral view), 35—female accessory sclerite. Scale bar: 0.1 mm.

FIGURES 23–26. Aedeagus of Paraphloeostiba (23–24: P. cooteri (paratype), 25–26: P. gayndahensis (Hong Kong)): 23, 25—parameral view, 24, 26—lateral view. Scale bar: 0.1 mm.