Histricostoma caucasicum (Redikortsev, 1936)

Modebadze, Naia, Martens, Jochen, Snegovaya, Nataly & Barjadze, Shalva, 2023, An annotated list of the Georgian harvestmen (Arachnida, Opiliones), Caucasiana 2, pp. 211-230 : 211

10. Histricostoma caucasicum (Redikortsev, 1936)

Nemastoma caucasicum Redikorzev, 1936: 37 (figs 7-8, 14, original description)

Nemastoma caucasicum - Roewer, 1951: 135 (fig. 65, redescription)

Nemastoma caucasicum - Mkheidze, 1964: 118 (mention)

Histricostoma (Histricostoma) caucasicum - Staręga, 1966: 394 (fig. 8, redescription)

Histricostoma caucasicum - Staręga, 1978: 201 (mention)

Histricostoma caucasicum - Martens, 2006: 192-195 (figs 21a-b, 26-27, redescription, distribution, elevational records, relationships)

Histricostoma caucasicum - Snegovaya, Pkhakadze and Intskirveli, 2014: 199 (mention)

Histricostoma caucasicum - Schönhofer, 2013: 30 (mention)

Histricostoma caucasicum - Snegovaya, 2013: 185 (mention)

Type locality.

Georgia, Abkhazia, Sokhumi Municipality, villages Ažary and Lata.

Occurrence data in Georgia.

Abkhazia • Sokhumi; leg. M. Kalichevsky, 10 August 1905 / 1 September 1905 ( Redikorzev 1936). • Mountain Apiancha, Tsebelda, Gudauta Municipality; 2500 m a.s.l.; leg. Y.V. Voronov, 9 September 1908 (Snegovaya et al. 2014). • Ritsa Lake, Gudauta Municipality; 950-1100 m a.s.l.; leg. S.I. Golovatch, 13/14 August 1986; Fagus , Abies , Picea , Acer etc. forest, litter, under bark and stones (CJM). • Pskhu, Sokhumi Municipality; 700-950 m a.s.l.; leg. S.I. Golovatch, 15/16 August 1986; Fagus , Quercus , Castanea etc. forest, litter, under bark and stones (CJM). Adjara • Batumi Botanical Garden, Kobuleti Municipality; leg. B. Pisarksi, 10 November 1963 ( Staręga 1978). • Zeraboseli, Kintrish State Reserve, Kobuleti Municipality; 450-600 m a.s.l.; leg. S.I. Golovatch and J. Martens, 1/3 June 1981 (CJM). • Zeraboseli, Kintrish State Reserve, Kobuleti Municipality; 600-800 m a.s.l.; leg. S.I. Golovatch and J. Martens, 2 June 1981 (CJM). • Batumi Botanical Garden; 20-150 m a.s.l.; leg. S.I. Golovatch and J. Martens, 5/7 June 1981 (CJM). • Batumi Botanical Garden; leg. S.I. Golovatch, 9 October 1981 (CJM). • 3 km W of Danisparauli, Khulo Municipality; leg. S.I. Golovatch and J. Martens, 10 October 1981; deciduous forest, litter (CJM). • Zeraboseli, Kintrish State Reserve, Kobuleti Municipality; 450-600 m a.s.l.; leg. S.I. Golovatch, 13 November 1981; deciduous forest, litter and under stones (CJM). • 6 km W of Khulo, Khulo Municpality; 800 m a.s.l.; leg. S.I. Golovatch, 11 November 1981; deciduous forest on rocky slope, litter (CJM). • 7 km NE of Batumi; 550 m a.s.l.; leg. V. Brachat and H. Meybohm, 9 June 2021 (CJM). • Chakvistavi, Kobuleti Municipality; 280 m a.s.l.; leg. V. Brachat and H. Meybohm, June 2021 (CJM). Samegrelo-Zemo Svaneti • 10 km N of Jvari, Tsalenjikha Municipality; 800 m a.s.l.; leg. S.I. Golovatch, 20/21 August 1986; Buxus , Fagus , Picea , Taxus etc. forest, litter (CJM). • Kherkhvashi, 40 km W of Mestia, Mestia Municipality; 1250-1700 m a.s.l.; leg S.I. Golovatch, 21 August/ 21 September 1986; Quercus , Fagus , Carpinus , Picea , Abies etc. forest, litter and bark (CJM). • Nenskra valley, Mestia Municipality; 800 m a.s.l.; leg. A.B. Ryvkin, 2 September 1986 (CJM). • Khaishi, Mestia Municipality; 600 m a.s.l.; leg. A.B. Ryvkin, 5 September 1986 (CJM). • Enguri valley, Mestia Municipality; 1000 m a.s.l.; leg. A.B. Ryvkin, 19 September 1986 (CJM). • W of Dizi, Mestia Municipality; leg. A.B Ryvkin, 19 September 1986 (CJM). • Mestia; 1500 m a.s.l.; leg. S.I. Golovatch, 16 September 1986 (CJM). • 40 km W of Mestia, Mestia Municipality; 1900-2200 m a.s.l; leg. S.I. Golovatch, 21 September 1986. • Mestia; 1910-1650 m a.s.l.; leg. J. and B. Martens, 22 July 2019; mixed forest (CJM). • Latpari Pass, Mestia Municipality; 1510-2100 m a.s.l.; leg. V. Brachat and H. Meybohm, 16 June 2021 (CJM). • 25 km N of Jvari, Tsalenjikha Municipality; 600 m a.s.l.; leg. J. and B. Martens, 20 July 2019; damp broad-leaved forest on a steep slope, ground moss covered (CJM). • Lakhamula, Mestia Municipality; 1110 m a.s.l.; leg. J. and B. Martens, 21 July 2019; open broad-leaved forest (CJM). Imereti • Sataplia State Reserve; 400 m a.s.l.; leg. S.I. Golovatch and J. Martens, 5 June 1981 (CJM). • between Tkibuli and Nikortsminda, Tkibuli Municipality; leg. S.I. Golovatch, 22 October 1981; Abies and Fagus forest, litter (CJM). • Mukhura, Tkibuli Municipality; 700-800 m a.s.l.; leg. S.I. Golovatch and K. Eskov, 7/9 May 1987; Castanea , Fagus , Carpinus etc., forest, litter, under bark and stones (CJM). • Villages Tsipa and Pona, Kharagauli Municipality; 980-1030 m a.s.l.; leg. J. and B. Martens, 16 July 2019; low Fagus orientalis forest, banks of a stream (CJM). Racha-Lechkhumi and Kvemo Svaneti • Timotesubani monastery, Borjomi Municipality; 1200 m a.s.l.; leg. J. and B. Martens, 15 July 2019; mixed forest along a stream (CJM). • Village Shkedi, Lentekhi Municipality; 720 m a.s.l.; leg. J. and B. Martens, 20 July 2019 (CJM). • S of Panaga, Lentekhi Municipality; 1250 m a.s.l.; leg. V. Brachat and H. Meybohm, 16 June 2021 (CJM). • Likheti, Ambrolauri Municipality; 1090 m a.s.l.; leg. V. Brachat and H. Meybohm, 17 June 2021 (CJM). Samtskhe-Javakheti • 15 km W of Adigheni, Adigeni Municipality; 1500-1700 m a.s.l.; leg. S.I. Golovatch, 14/15 May 1983; Abies , Picea , Fagus , Acer etc. forest, litter, logs, under stones (CJM). • 8 km SE of Akhaldaba, Borjomi Municipality; 1000 m a.s.l.; leg. S.I. Golovatch, 12 May 1983; Nedzura River valley, Picea , Carpinus and Fagus forest, litter, logs (CJM). • Borjomi-Kharagauli National Park; 800-900 m a.s.l.; leg. S.I. Golovatch, 12/16 May 1983; Picea , Fagus and Carpinus forest, litter, logs, under stones (CJM). • Borjomi; 960 m a.s.l.; leg. J. and B. Martens, 17 July 2019; park and forest along the Mtkvari river and small stream, mixed broad-leaved forest (CJM). Shida Kartli • Surami, Khashuri Municipality; 850 m a.s.l.; leg. S.I. Golovatch and K. Eskov, 7 May 1987 (CJM). • Mariamjvari Nature Reserve, Sagarejo Municipality; 1150-1250 m a.s.l.; leg. S.I. Golovatch and K. Eskov, 13/14 May 1987; Fagus , Carpinus , Acer , Pinus etc. forest, litter, under bark and stones (CJM). • Algeti National Park, Tetritskaro Municipality; 1400-1450 m a.s.l.; leg. S.I. Golovatch and Eskov, 16/18 May 1987; Fagus , Picea , Acer etc. forest, litter and under bark (CJM). • SW of Patara Dmanisi, Bolnisi Municipality; 1000-1050 m a.s.l.; leg. S.I. Golovatch and Eskov, 21 May 1987; Quercus , Fagus , Acer etc. forest, litter (CJM). • S of Khorisari Pass, Kazbegi Municipality; 1500 m a.s.l.; leg. S.I. Golovatch, 17 October 1987; Fagus , Betula etc. forest, litter (CJM). • SW of Nikortsminda, Ambrolauri Municipality, 1050 m a.s.l.; leg. S.I. Golovatch, 19 October 1987; Fagus , Abies , Acer etc. forest (CJM). • Kvishketi, Holy Mountain above the village; 1200 m a.s.l.; leg. J. and B. Martens, 15 July 2019; Fagus orientalis forest (CJM). • Khashuri Municipality; 830 m a.s.l.; leg. J. and B. Martens, 16 July 2019; mixed forest, mainly Picea orientalis , on stream banks (CJM). Mtskheta-Mtianeti • Mtskheta; leg. Roszkowski, 3 May 1918 ( Staręga 1978). Kakheti • Lagodekhi Nature Reserve, Lagodekhi Municipality; 600-700 m a.s.l.; leg. S.I. Golovatch, 5/6 May 1983; Fagus , Fraxinus , Acer etc. forest, litter, logs, under stones (CJM).• N of Kvareli; 700-750 m a.s.l.; leg. S.I. Golovatch and K. Eskov, 4 May 1987; Fagus , Carpinus , Quercus etc. forest, litter and under bark (CJM). • 20 km N of Akhmeta, Batsaro State Reserve; 800-850 m a.s.l.; leg. S.I. Golovatch and K. Eskov, 5/6 May 1987; Fagus , Castanea forest, litter (CJM). • Dzveli Shuamta’s Monastery, Telavi Municipality; leg. J. and B. Martens, 28 July 2019; old Fagus orientalis forest (CJM). • Gombori Pass, Sagarejo Municipality; 1500-1600 m a.s.l.; leg. J. and B. Martens, 28 July 2019; bushes along the stream (CJM). • Birkiani, Akhmeta Municipality; 930-1100 m a.s.l.; leg. J. and B. Martens, 31 July 2019; side valley, mixed broad-leaved forest, river banks (CJM).

Global distribution.

Endemic to the Caucasian ecoregion: Georgia, Azerbaijan, Russian Caucasus, Armenia, Turkey ( Staręga 1978; Martens 2006; Schönhofer 2013; Snegovaya 2013).

Remarks.

In Georgia the species is widely distributed and it is the most common species of the small litter-inhabiting nemastomatids. In Georgia, its known elevational distribution spans from 280 m to 2100 m (CJM); for records beyond Georgia see Martens (2006).