Coenosia projecta (Malloch, 1934)

Patitucci, Luciano Damian, Mulieri, Pablo Ricardo, Couri, Marcia Souto & Dominguez, Martha Cecilia, 2023, Phylogeny of the old and fragmented genus Austrocoenosia Malloch reveals new evidences on the morphology and evolution of the genera Coenosia Meigen and Neodexiopsis Malloch (Diptera: Muscidae), Arthropod Systematics & amp; Phylogeny 81, pp. 611-653 : 611

Coenosia projecta (Malloch, 1934)

Male (Fig. 26A).

Length. Body: 2.80-3.04 mm, wing: 2.25-2.48 mm. Head: (Fig. 26B ). Frons at vertex about one third of the head width. Frons, fronto-orbital plate, parafacial and gena black with grey pollinosity; 3-4 pairs of frontal setae. Frons longer than wide, with frontal triangle long, light grey, reaching lunula. Gena narrow, narrower than the width of postpedicel. Fronto-orbital plate with 3-5 little setulae, close to parafacialia. Antenna black, apical angle of postpedicel acute; in lateral view inserted over the mid-level of the eye; arista with its longest microtrichia shorter than its basal diameter. Thorax: (Fig. 26C ). Black with grey pollinosity, with three black vitta at dorsocentral and acrostichal rows of setae; anterior and posterior spiracles brown. Chaetotaxy: acr s short and irregular, the anterior presutural acr s longer than the anterior presutural dorsocentral seta (eaDC); dorsocentrals 1+3, eaDC is less than one third of the aDC. Katepisternum with 3-4 setulae. Wing: Tegula black. Both calypters whitish hyaline with white margins; halter yellow. Legs: Black with grey pollinosity, apex of femora yellow. Fore femur with a row of strong pd, a row of strong pv setae, and a row of av setae at basal third. Mid femur with 5-6 setae on anterior surface on basal half, 3-5 pv setae on basal third and a row of av setae, 2 preapical setae on pd to posterior surface; mid tibia with one ad and one pd median setae, with similar length. ad seta placed apical than pd seta. Hind femur with a complete row of ad and a row of av setae on apical half, with 3-4 pv setae on basal third. Similar size of claws and pulvilli of the three legs. Abdomen: (Fig. 26D ). Black, with grey pruinosity, tergites 3-5 with two dark brown spots. Tergite 6 not visible in dorsal view. Sternite 5 broader than long, apical margin concave with a “U” shape, with a depressed area at the base, and without membrane; setae concentrated on the lobes and some long and strong on apical margin (Fig. 26E ). Terminalia: Cercus curved in lateral view, keels placed from distal to basal margin, apical margin straight. Surstylus longer than broad curved at apex, sclerotized (Fig. 26F, G ). Hypandrium tubular, wider than long, distal extreme not exposing the phapod, and without spines on ventral surface. Aedeagus with pregt developed and sclerotized; pgt developed; epiphalus slightly sclerotized, and distiph tubular, sclerotized at base, and without hairs on ventral surface (Fig. 26G, H ).

Female.

Unknown.

Type material.

Holotype. Male; BMNH [pinned, in good condition]. Original labels: "♂ [handwritten] / Holo- / type [printed]" on white circular paper, red frame; "Argentina: / Terr. Rio Negro. / F.& M. Edwards / B.M. 1927-63." printed on white paper; "R. Pichileufu / 24.x.1926." printed on white paper; " Austrocoenosia / Austrocoenosia projecta / Type [handwritten] / det. JRMALLOCH [printed]" on white paper, black frame. https://oumnh.ox.ac.uk/collections-online#/item/oum-catalogue-381555.

Additional material examined.

ARGENTINA - Chubut province • 3 males, PNLA, Seccional Lago Verde ; -42.718803; -71.727470; 538 m a.s.l.; Oct 2014; Mulieri & Patitucci leg.; MACN .

Distribution (Fig. 27E).

ARGENTINA: Chubut (new record), Río Negro.

Remarks.

Coenosia projecta was described by Malloch (1934) with one male specimen from Río Negro, Argentina. Malloch (1934: 224 Fig. 38b) illustrated some striking long and fine hairs on the ventral surface of the hind femur, but did not mention anything in the description. We did not observe these hairs in the images of the type specimen or in the specimens collected, so we assume that this was a mistake in the original drawing.

Figures

Figure 26. Coenosia projecta, male. A Lateral view, B head, frontal view, C thorax, dorsal view, D abdomen, dorsal view, E fifth sternite, F cercus and surstylus, dorsal view, G cercus, surstylus and phallic complex, lateral view, H phallic complex, lateral view. Scale bar: 1 mm (A), 0.5 mm (B-D), 0.2 mm (E-G), 0.05 mm (H).