Ancognatha ustulata (Burmeister, 1847)

Moore, Matthew R., Cave, Ronald D. & Branham, Marc A., 2018, Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini), ZooKeys 745, pp. 101-378 : 116-117

Ancognatha ustulata (Burmeister, 1847)

Cyclocephala ustulata Burmeister, 1847: 39 [original combination].

Ancognatha ustulata (Burmeister) [new combination by Bates 1888: 297].

syn. Ancognatha ustulata ustulatoides Höhne, 1922d: 373-374 [original combination].

Ancognatha ustulata var. ustulatoides Höhne [new infrasubspecific status by Arrow 1937b: 6].

Ancognatha ustulata ab. ustulatoides Höhne [revised infrasubspecific status by Endrődi 1985a: 162].

Types.

Lectotype ♂ of C. ustulata at MLUH ( Endrődi 1966). Lectotype ♂ of A. ustulata ustulatoides at ZMHB ( Endrődi 1966).

Distribution.

COLOMBIA: Antioquia, Bogotá D. C., Boyacá, Caldas, Cauca, Chocó, Cundinamarca, Tolima, Valle del Cauca. ECUADOR. PERU: Pasco. VENEZUELA: Mérida.

References.

Dejean 1833, 1836b, Sturm 1843, Burmeister 1847, Höhne 1922d, Arrow 1937b, Blackwelder 1944, Apolinar Maria 1946, Pike et al. 1976, Endrődi 1966, 1985a, Restrepo et al. 2003, Neita-Moreno and Morón 2008, Gasca-Álvarez and Amat-García 2010, Krajcik 2005, 2012, Breeschoten et al. 2013, López-García et al. 2015, Figueroa and Ratcliffe 2016.

Remarks.

Ancognatha ustulata ustulatoides was a validly described subspecies ( Höhne 1922d). The subspecies was considered an infrasubspecific entity after Arrow (1937b) and was referred to as an “ab.” or “var.” (e.g., see Endrődi 1966, 1985a). The name has not been clearly synonymized with A. ustulata (Burmeister), but was listed as a synonym by Krajcik (2005). Endrődi (1966, 1985a) reported A. ustulata from Mexico and Panama, but major faunistic studies have not found additional specimens from these countries ( Ratcliffe 2003, Ratcliffe et al. 2013).