Asiobaccha tripartita ( Walker, 1861 ), Mengual, Ximo, 2016

Mengual, Ximo, 2016, A taxonomic revision of the genus Asiobaccha Violovitsh (Diptera: Syrphidae), Journal of Natural History 50, pp. 2585-2645: 2632-2634

publication ID

http://doi.org/ 10.1080/00222933.2016.1206634

publication LSID

lsid:zoobank.org:pub:6B627B0F-9440-47F1-90F4-9AF4C7308A99

DOI

http://doi.org/10.5281/zenodo.4333765

persistent identifier

http://treatment.plazi.org/id/B23987DC-FFA9-FFF0-5E30-D4A2FCE9FB1A

treatment provided by

Carolina

scientific name

Asiobaccha tripartita ( Walker, 1861 )
status

comb. nov.

Asiobaccha tripartita ( Walker, 1861)   comb. nov.

( Figures 11c, d, f View Figure 11 , 14d View Figure 14 )

Baccha tripartita Walker, 1861: 285   . Holotype: ♂, BMNH. Type locality: Indonesia: Misool Island. Schiner 1868: 344; Bigot 1892: 166; Wulp 1896: 121; Kertész 1910: 157.

Baccha flavipes Doesburg, 1959: 233   . Syn. nov.

Baccha sulica Austen, 1893: 144   (in part). Syn. nov. Wulp 1896: 121; Kertész 1910: 157.

Baccha (Allobaccha) sulica   of Knutson et al. 1975: 323 (in part).

Baccha (Allobaccha) tripartita   of Knutson et al. 1975: 323.

Baccha papuana Hull   , unpublished of Greve and Ismay 1983: 74.

Episyrphus (Asiobaccha) sulica   of Thompson and Vockeroth 1989: 443 (in part).

Episyrphus (Asiobaccha) tripartita   of Thompson and Vockeroth 1989: 443.

Allobaccha flavipes   of Thompson and Vockeroth 1989: 441.

Allobaccha papuana   (Hull, unpublished name) of Rojo et al. 2003: 20.

Differential diagnosis

This species has a yellow face ( Figure 11f View Figure 11 ), the alula is bare, narrower than costal cell, the wing is partly bare basally ( Figure 14d View Figure 14 ), and it does not have a mesonotal fringe. The abdominal pattern is identical to that of two other species, A. bicolor   and A. loriae   , with tergum 4 entirely black ( Figure 11c, d View Figure 11 ), but differs by the wing microtrichia as stated in the key. Asiobaccha bicolor   and A. tripartita   differ from A. loriae   by having the wing bare basally, easy to distinguish because both always have the cell CuP and anal lobe partly bare basally ( Figure 14c, d View Figure 14 ); sometimes A. loriae   specimens may have very small bare areas in the cell BM but the cell CuP and the anal lobe are always microtrichose. On the other hand, A. tripartita   has a more extensive microtrichia and bare basal areas of the wing are on basal 1/3 – 1/2 cell BM only ( Figure 14d View Figure 14 ), while A. bicolor   has the cell R bare on basal 1/3 – 1/2 and the cell BM bare on basal 1/2 or more ( Figure 14c View Figure 14 ).

Variation. Wing microtrichia is a bit less variable in this species: the bare area of cell BM varies between basal 1/3 to basal 1/2. Furthermore, a few specimens have costal cell bare very basally. The specimen from New Ireland has the scutum entirely yelloworange. Wing colouration varies from very dark to almost hyaline.

Length (N = 4). Body, 11.0 – 13.0 (12.3) mm; wing, 10.0 – 12.0 (11.1) mm.

Biology

Greve and Ismay (1983) mentioned one species of Baccha   with the following information: ‘ Ex Tiracola   (Lep., Noctuidae   ) larva, Brevaturu, N.P., August, 1971 ’. The species mentioned is Baccha   ? [sic] papuana Hull. This   record is compiled by Rojo et al. (2003) but with a different name, Allobaccha papuana   (Hull, unpublished name).

In the opinion of this author, this species might refer to the unpublished name of Baccha papuana Hull   that appears in two specimens from Mafulu kept at the BMNH and CNC, although there is no information in the original labels about Tiracola   .

After studying the work of Greve and Ismay (1983), this citation may refer to a larva of a syrphid species parasiting a larva of Tiracola Moore, 1881   ( Lepidoptera   : Noctuidae   ) from Barevaturu N.P. in Papua New Guinea. However, there is no species of Asiobaccha   or Baccha   that parasitises butterfly larvae. Consequently, this record by Greve and Ismay (1983) may not refer to a species of Baccha   , Allobaccha   or Asiobaccha   , or, on the other hand, it might indicate that the larva of Baccha   was preying on the Tiracola   larva and the adult emerged from a puparium.

Geographical distribution

Kai Islands, Misool Island, New Guinea Island, New Ireland Island and Woodlark Island.

Type locality

Indonesia: West Papua Province, Raja Ampat Regency, Misool Island (Mysol), between 01°40 ’ and 02°04 ’ S, and between 129°43 ’ and 130°26 ’ E GoogleMaps   .

Material examined

Type material. Holotype, male, deposited in The Natural History Museum, formerly British Museum (Natural History) (London, United Kingdom) and labelled: ‘W’ [round, handwritten] ‘Mysol. // Wallace. ’ ‘ Baccha   // tripartita   // Wlk. ’ [on the reverse of previous label, handwritten] ‘tripartita’ [handwritten] ‘ HOLOTYPE // Asiobaccha   // tripartita   // (Walker) // det. X. Mengual 20 14’ [red].

Type material of Baccha sulica Austen, 1893   . Syntype, female, deposited in The Natural History Museum (London, UK) and labelled: ‘ SYN- // TYPE ’ [round, blue margin] ‘ Mysol ’ [round, light blue] ‘ hl // 124 ’ [on the reverse of previous label, handwritten] ‘Mysol. // (Wallace.) // 61.124. ’ ‘ Baccha   // sulica   // Aust. ’ [on the reverse of previous label, handwritten].

Type material of Baccha flavipes Doesburg, 1959   . Holotype, female, deposited in the Naturalis Biodiversity Center (Leiden, The Netherlands) and labelled: ‘ HOLOTYPE ’ [yellow] ‘G.den Hoed // Ifar 12-57’ [handwritten] ‘ Museum Leiden // Collectie // Van Doesburg // rec. 1973 ’ ‘ Baccha   ♀ // flavipes Dobg.   // det. v. Doesburg ’.

Nontype material. INDONESIA: Maluku Prov.   , Great Kai Island , Mountain Daab, April 1922, T. Mortensen [2♀, ZMUC]; Papua Prov   ., 11 km SE of Oerberfaren, Bodem , 100 m, 7 – 17 July 1959, T   .C. Maa [♀, BPBM]   ;..., Wataikwa River , August 1910, A   .F.R. Wollaston 1911 – 229 [1♀, BMNH]   ;..., Araucaria River , SW von Bernhard Camp, 800 m, 6 January 1939, L   .J. Toxopeus [1♀, RMNH]   . PAPUA NEW GUINEA: Central Prov   ., Owen Stanley Range , Goilala, Loloipa, 1 – 15 February 1958, W   .W. Brandt [1♂ 1♀, BPBM; 1♂, ZFMK, ZFMKDIP 00011948 View Materials ]   ;..., Goilala , Tapini, 975 m, 16 – 25 November 1957, W   .W. Brandt [1♂, BPBM]; Central Prov   ., Mafulu, 4000 ft., January 1934, L.E. Cheesman [1♀, BMNH; 1♀, CNC]; West Sepik Prov   ., Telefomin District , Eliptamin Valley, 1200 – 1350 m, 19 – 30 June 1959, W   .W. Brandt [1♀, BPBM]; Madang Prov   ., Finisterre Range , Saidor, Sibog village, 27 May – 5 June 1958, W   .W. Brandt [1♀, BPBM]; Jiwaka Prov   ., Upper Jimi valley , Tsenga, 1200 m, 15 July 1955, J   .L. Gressitt, light trap [1♂, BPBM]; Morobe Prov   ., Bulolo Gorge , 800 m, 17 January 1962, G   . Monteith [1♂, BPBM]   ;..., 6 miles NW Lae, Rain forest , 15 m, 9 July 1957, D   .E. Hardy [1♂, ZFMK, ZFMKDIP 00011952 View Materials ]   ;..., Bulolo , Manki Logging area, 2 June 1983, H   . Roberts [1♂, BMNH] (specimen photographed)   ;..., Wau , 6 – 14 March 1974, H   . Hippa, P.T. Lehtinen [1♀, MZH]   ;..., Wau , April 1968, M   . Sedlacek [1sp., BPBM]   ;..., Wau, Nakata Range, 4700 ft., 9 June 1968, J.W. Boyes [1♂, CNC]; Milne Bay Prov   ., Woodlark Island , Kulumadau Hill, 4 – 9 March 1957, W   .W. Brandt [1♂ 1♀, BPBM; 1♀, ZFMK, ZFMKDIP 00011950 View Materials ]   ;..., 16 March 1957,... [1♀, ZFMK, ZFMKDIP 00011953 View Materials ]; New Ireland Prov   ., Danu , Kalili Bay, 3 April 1962, Noona Dan Exp   . 61 – 62 [1♂, ZMUC]   ;..., 30 April 1962,... [1♀, ZMUC]   .

Remarks

Austen (1893) described Baccha sulica   based on two female specimens, but only the female from Sula Island had Walker ’ s handwriting. Austen (1893) stated that he found the type of this species, but he could not find if it was already described. In my opinion, he decided to describe a new species based on Walker ’ s handwriting. The most interesting fact is that Austen (1893) used the same argument to separate sulica   from Baccha moluccana   as to separate bicolor   from moluccana   , but he did not compare his two new species. In the BMNH there are the two syntypes of Baccha sulica   , but one belongs to A. bicolor   (specimen from Sula Island and Walker ’ s handwriting) and the other syntype belongs to A. tripartita   (specimen from Misool).

Specimens from the New Ireland at ZMUC have a manuscript name given by P.H. van Doesburg, Baccha pleuralis   , never published. Specimens from Mafulu have a manuscript name given by F. M. Hull, Baccha papuana   , again never published. None of these three names is available.

As already mentioned, A. tripartita   might belong to a widespread taxon and might include also A. loriae   and A. bicolor   (see Remarks under A. bicolor   and A. loriae   ). There are specimens of A. tripartita   and A. bicolor   from the same localities, but, more interestingly, specimens of bicolor   and tripartita   from New Ireland Island have an orange scutum. Both females from Great Kai Island also have the scutum orange.

ZMUC

Zoological Museum, University of Copenhagen

BPBM

Bishop Museum

RMNH

National Museum of Natural History, Naturalis

ZFMK

Zoologisches Forschungsmuseum Alexander Koenig

CNC

Canadian National Collection of Insects, Arachnids, and Nematodes

MZH

Finnish Museum of Natural History

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Diptera

Family

Diptera (awaiting allocation)

Genus

Asiobaccha

Loc

Asiobaccha tripartita ( Walker, 1861 )

Mengual, Ximo 2016
2016
Loc

Allobaccha papuana

Rojo S & Gilbert F & Marcos-Garcia MA & Nieto JM & Mier MP 2003: 20
2003
Loc

Episyrphus (Asiobaccha) sulica

Thompson FC & Vockeroth JR 1989: 443
1989
Loc

Episyrphus (Asiobaccha) tripartita

Thompson FC & Vockeroth JR 1989: 443
1989
Loc

Allobaccha flavipes

Thompson FC & Vockeroth JR 1989: 441
1989
Loc

Baccha papuana

Greve JE & Ismay JW 1983: 74
1983
Loc

Baccha (Allobaccha) sulica

Knutson LV & Thompson FC & Vockeroth JR 1975: 323
1975
Loc

Baccha (Allobaccha) tripartita

Knutson LV & Thompson FC & Vockeroth JR 1975: 323
1975
Loc

Baccha flavipes

Doesburg PJ 1959: 233
1959
Loc

Baccha sulica

Kertesz K 1910: 157
van der Wulp FM 1896: 121
Austen EE 1893: 144
1893
Loc

Baccha tripartita

Kertesz K 1910: 157
van der Wulp FM 1896: 121
Bigot JMF 1892: 166
Schiner IR 1868: 344
Walker F 1861: 285
1861