Glyphiulus javanicus, Carl, 1911

Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul & Spiegel, Didier Van Den, 2007, Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 2: the javanicus-group, Zoosystema 29 (3), pp. 417-456 : 420-422

The javanicus -group

REMARKS

The javanicus -group can be characterised as follows:

1. Male legs 1 are normal to reduced, never hypertrophied; telopodites often complete or nearly so, even when reduced in size, usually multisegmented; coxae or coxosternum with a pair of small but evident, central, medially nearly always contiguous, sometimes partly even fused outgrowths or processes which are never coalesced completely into a strong and long sickle-shaped structure (NB: a few species known only from female material are assigned here only provisionally).

2. Male legs virtually normal, not enlarged.

3. Typical carinotaxy patterns of the collum are I-III(IV, V)+P+M+P+(V,IV)III-I or, as in the granulatus -group, I-VI+7a+pc+ma+pc+7a+VI-I, or something close. The patterns are often different, often due to crest reduction (especially anteriorly), but a median crest is always visible at least near the caudal margin.

4. The typical carinotaxy pattern of the metaterga is 2(1)/2(1)+I/i+3/3+I/i+2(1), thus reflecting the fact that all or most of the crests are usually divided transversely into two halves. In contrast to species of the granulatus -group, the median crest is never doubled anteriorly, but sometimes it is more or less strongly reduced. If the pattern is different, it can usually be readily derived from the typical one (or vice versa), i.e. the lateral crests are reduced to one or the crests are undivided (see also below), etc.

5. The anterior gonopods are usually reduced to a platelike coxosternum with moveable, lateral, 1-segmented, often curved telopodites (shared with some other species groups).

6. The posterior gonopods are usually strongly compressed, often showing a subflagelliform, sometimes plumose, distal flagellum (shared with some other species groups).

7. The pleural flaps behind the gonopod opening on male segment 7 usually form a clear, transverse, ventral ridge (shared with some other species groups).

The remaining known cambalopsids are considered to belong to other species groups, partly already treated ( Golovatch et al. 2007) and partly to be treated elsewhere.

Below, a description is given of the available material, followed by a key to all of the constituent species of the javanicus- group, and a brief analysis of their variation and distribution.