Glyphiulus subobliteratus, Golovatch & Geoffroy & Mauriès & Spiegel, 2007

Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul & Spiegel, Didier Van Den, 2007, Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 2: the javanicus-group, Zoosystema 29 (3), pp. 417-456 : 424-426

publication ID

https://doi.org/ 10.5281/zenodo.5393630

persistent identifier

https://treatment.plazi.org/id/B36487D8-FFC6-D531-D10A-FEA6FB09F95A

treatment provided by

Marcus

scientific name

Glyphiulus subobliteratus
status

sp. nov.

Glyphiulus subobliteratus View in CoL n. sp.

( Figs 3-5 View FIG View FIG View FIG )

TYPE MATERIAL. — China. Yunnan Prov., Shilin County, 85 km SE of Kunming, Shilin (Stone Forest), Zi Yung Dong Cave , 1.X.1988, leg. P. Beron, holotype ♂ ( NMNHS), paratypes 5 ♂♂, 10 ♀♀, 9 juvs ( NMNHS) ; 1 ♀ ( SEM) ; 2 ♂♂, 2 ♀♀ ( MNHN GA 045 View Materials ) ; 1♂, 1 ♀ ( ZMUM) ; 1 ♂, 1 ♀ ( ZMUC) ; 1 ♂, 1 ♀ ( MCSNV) .

ETYMOLOGY. — To emphasize the apparent resemblance to G. obliteratus n. sp.

DIAGNOSIS. — Differs from the most closely related species G. obliteratus n. sp. and Glyphiulus zorzini by the pigmented body, the collum undulate caudally (more strongly so than in G. obliteratus n. sp. but less so than in G. zorzini ), the mostly higher and subdivided metatergal crests, some details of gonopod structure, etc. (see also key below).

DESCRIPTION

Holotype with 53p+2a+ T, about 36 mm long and 2.0 mm wide; adult paratypes (♂, ♀) with 42-56p+4- 1a+ T, length 27-40 mm, width 1.6-2.0 mm.

Coloration marbled pale to dark grey-brown, especially infuscate due to metatergal crests; venter, antennae and legs yellowish. Ocelli black-brown, arranged in two transverse rows, seven or eight in each patch.

Antennae ( Figs 3A View FIG ; 4C View FIG ; 5A View FIG ), gnathochilarium (mentum divided, n = 7) ( Fig. 5B View FIG ), segment shape ( Fig. 4A View FIG ), epiproct ( Fig. 3E, F View FIG ), male legs 2 and 3 ( Fig. 5E, F View FIG ), pleurosternal crests on segments 2-4 ( Fig. 3A View FIG ), accessory spine at base of claw ( Fig. 4D View FIG ) and posterior gonopods ( Fig. 5 View FIG H-K) generally as in G. obliteratus n. sp., but crests/undulations on collum low and incomplete, formula 1a+2p+3a-4a+ma+4a- 3a+2p+1a ( Fig. 3A, B View FIG ); crests on subsequent segments higher (even adding another lateral crest below ozoporiferous one), mostly indistinctly subdivided (except for several anterior- and posteriormost segments); carinotaxy pattern 2/2+I/i+3/3+I/i+2/2, with anterior parts smaller than posterior ones ( Fig. 3 View FIG ); legs shorter than body diameter ( Fig. 4A View FIG ); hypoproct with several scattered setae ( Fig. 4B View FIG ); male leg 1 with particularly strongly developed paramedian coxal outgrowths and 5-segmented, though slightly reduced, telopodites ( Fig. 5C, D View FIG ); notch on each half-anterior gonopod less deep ( Fig. 5G View FIG ), etc.

Head width = segment 2 = 6 <collum = 7(8) = midbody segments> segment 3> 4> 5; body first gently but then abruptly tapering toward telson. Postcollar constriction evident ( Fig. 3B View FIG ).

Ozoporiferous cones round, wider than high ( Fig. 4A View FIG ).

Penes broad, rounded, each with two to four strong setae distolaterally ( Fig. 5E View FIG ).

Anterior gonopods ( Fig. 5G View FIG ) with both digitiform (d) and subsecuriform (s) processes lower than in G. obliteratus n. sp.

REMARKS

The sample contains three haplopodous juveniles with 28p+2a+T, each with the ozoporiferous tubercles fully developed only on segments 5 and 6.

In the absence of any troglomorphic traits, this species can only be considered troglophilic at most.

MNHN

Museum National d'Histoire Naturelle

ZMUM

Zoological Museum, University of Amoy

ZMUC

Zoological Museum, University of Copenhagen

T

Tavera, Department of Geology and Geophysics

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