Glyphiulus paracostulifer, Golovatch & Geoffroy & Mauriès & Spiegel, 2007

Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul & Spiegel, Didier Van Den, 2007, Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 2: the javanicus-group, Zoosystema 29 (3), pp. 417-456 : 442-446

publication ID	https://doi.org/ 10.5281/zenodo.5393630
persistent identifier	https://treatment.plazi.org/id/B36487D8-FFD8-D525-D108-FA5BFEC9F9F8
treatment provided by	Marcus
scientific name	Glyphiulus paracostulifer
status	sp. nov.

Treatment

Glyphiulus paracostulifer View in CoL n. sp.

( Figs 22-24 View FIG View FIG View FIG )

TYPE MATERIAL. — China. Guizhou Prov., Qianxy, Hong Lin Town, Lao Hu Dong Cave , 21.XI.2003, leg. L. Latella & S. Meggiorini, holotype ♂ ( MNHN GA 052 View Materials ) ; paratypes 1 ♀ ( MNHN GA 052 View Materials ) ; 1 ♀ ( ZMUM) ; 1 ♀ ( MCSNV / SEM) .

ETYMOLOGY. — To emphasize the close resemblance to G. costulifer n. sp.

DIAGNOSIS. — Differs from congeners in the relatively large size, the completely unpigmented tegument and ocelli, the undivided tergal crests, coupled with certain details of leg and gonopod structure (see also key below).

DESCRIPTION

Holotype with 65p+1a+T, about 43 mm long and 2.0 mm wide; adult paratypes with 52p+2a+T, 55p+3a+T or 57p+1a+T, all about 36 mm long and 2.0 mm wide.

Coloration rather uniformly pale yellowish to brownish, only some ozoporiferous cones a little darker; ocelli unpigmented, a few barely discernible per eye patch.

Gnathochilarium (mentum divided, n = 1), postcollar constriction ( Fig. 22B View FIG ), segment’s texture and cross-section ( Fig. 22 View FIG A-E), most of telson ( Fig. 22E, F View FIG ), ventral flaps of male segment 7, male legs 2 and 3 ( Fig. 24C, D View FIG ), posterior gonopods ( Fig. 24G View FIG ) generally as in G. subcostulifer n. sp., but antennae slightly more elongate and less rich in sensilla ( Figs 22A View FIG ; 23D View FIG ; 24A View FIG ); collum ( Fig. 22A, B View FIG ) with a nearly typical carinotaxy formula (I-III+4c+5a+pc+ma+pc+5a+4c+III- I), crests evident; all tergal crests ( Figs 22 View FIG A-E; 23A) on subsequent body segments higher, especially so until about segment 9 or 10, virtually not divided (2+I/i+3+I/i+2), ozoporiferous cones only barely lower than wide; epiproct with an evident dorsal rib ( Fig. 22E View FIG ); legs about as long as body diameter ( Fig. 23A, B View FIG ), at most with only a vestigial spine at claw base ( Figs 23C View FIG ; 24E View FIG ); male leg 1 with reduced 5-segmented telopodites and large paramedian coxal processes ( Fig. 24B View FIG ); each anterior gonopod plate with a high distomedial process ( Fig. 24F View FIG ).

REMARKS

Based on several troglomorphic traits, such as the unpigmented body and ocelli, as well as the somewhat elongated antennae and legs, this species seems to be troglobitic.