Glyphiulus percostulifer, Golovatch & Geoffroy & Mauriès & Spiegel, 2007

Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul & Spiegel, Didier Van Den, 2007, Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 2: the javanicus-group, Zoosystema 29 (3), pp. 417-456 : 441-442

publication ID	https://doi.org/ 10.5281/zenodo.5393630
DOI	https://doi.org/10.5281/zenodo.5469542
persistent identifier	https://treatment.plazi.org/id/B36487D8-FFD9-D521-D2D5-FD42FC5AF95A
treatment provided by	Marcus
scientific name	Glyphiulus percostulifer
status	sp. nov.

Treatment

Glyphiulus percostulifer View in CoL n. sp.

( Figs 20 View FIG ; 21 View FIG )

TYPE MATERIAL. — Laos. Khammouan Prov., Ban Nakok (= Nakhok), Tham Thê , 104.49°E, 17.90°N, cave, hand collection, 11.II.1998, leg. A. Bedos & L. Deharveng (LAO-016), holotype ♂ ( MNHN GA 051 About MNHN ) GoogleMaps ; paratypes 1 ♂, 1 ♀, 1 subad. ♂ ( MNHN GA 051 About MNHN ) GoogleMaps ; 1 ♀ ( ZMUM / SEM) .

ETYMOLOGY. — To emphasize the very close resemblance to G. costulifer n. sp., but with more strongly developed tergal crests.

DIAGNOSIS. — Differs from congeners in the small size, the completely unpigmented tegument and ocelli, coupled with certain details of leg and gonopod structure (see also key below).

DESCRIPTION

Holotype with 34p+2a+T, about 14 mm long and 1.0 mm wide; adult paratypes with 36p+2a+T, about 14 mm long and 1.0 mm wide (♂), with 40p+1a+T or 38p+1a-T, about 17 mm long and 1.0 mm wide (♀); subadult ♂ with 31p+2a+T.

Coloration uniformly pallid, ocelli not visible ( Fig. 20A View FIG ).

Postcollar constriction, dorsal crests and carinotaxy pattern on postcollar terga ( Fig. 20 View FIG A-E), segment’s texture and cross-section ( Fig. 20 View FIG B-D), paraprocts and hypoproct ( Fig. 20F View FIG ), ventral flaps of male segment 7, male legs 2 and 3 ( Fig. 21D, E View FIG ), penes ( Fig. 21D View FIG ) generally as in G. costulifer n. sp., but antennae elongate, antennomeres 5-7 richer in sensilla ( Fig. 21A View FIG ), gnathochilarium (mentum divided, n = 2) ( Fig. 21B View FIG ) slightly even more oligotrichous; collum ( Fig. 20A View FIG ) with a nearly typical carinotaxy formula (I-IV+5a+pc+ma+pc+5a+IV-I); all tergal crests ( Fig. 20 View FIG A-E) on subsequent body segments higher, subdivided (2/2+I/i+3/3+I/i+2/2), ozoporiferous cones about as high as wide; epiproct ( Fig. 20E, F View FIG ) upturned and broadly rounded in caudal part, with one median and two paramedian crests; legs slightly longer than body diameter ( Fig. 20B View FIG ), always with an accessory filament/spine at base of claw, which is usually slightly more than half as long as claw ( Fig. 21 View FIG D-F); male leg 1 with normal, 5-segmented telopodites and large, paramedian, coxal processes ( Fig. 21C View FIG ); each anterior gonopod plate with a high and subsecuriform distomedial process ( Fig. 21G View FIG ); posterior gonopods with distally slightly plumose flagella ( Fig. 21H View FIG ).

REMARKS

Based on several troglomorphic traits, such as the completely unpigmented body and ocelli, as well as the elongated antennae and legs, this species seems to be troglobitic, although the small size may also be evidence of its being an inhabitant of the MSS (“milieu souterrain superficiel”).