Glyphiulus intermedius, Golovatch & Geoffroy & Mauriès & Spiegel, 2007

Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul & Spiegel, Didier Van Den, 2007, Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 2: the javanicus-group, Zoosystema 29 (3), pp. 417-456 : 446-449

publication ID	https://doi.org/ 10.5281/zenodo.5393630
DOI	https://doi.org/10.5281/zenodo.5469544
persistent identifier	https://treatment.plazi.org/id/B36487D8-FFDC-D518-D2D2-F9FAFBC1FA99
treatment provided by	Marcus
scientific name	Glyphiulus intermedius
status	sp. nov.

Treatment

Glyphiulus intermedius View in CoL n. sp.

( Figs 25 View FIG ; 26 View FIG )

TYPE MATERIAL. — China. Sichuan Prov., Chengdu County, Huan Long Dong Cave ( No. 593), 26.VIII.1999, leg. J. & B. Lips, holotype ♂ ( MNHN GA 053 About MNHN ) ; paratypes 1 ♂, 2 ♀♀, 1 juv ( MNHN GA 053 About MNHN ) ; 1 ♀ ( SEM) ; 1 ♀ ( ZMUM) .

ETYMOLOGY. — To emphasize a somewhat intermediate position between Podoglyphiulus species and the remaining species of the javanicus -group.

DIAGNOSIS. — Differs from congeners in the particularly long ozoporiferous tubercles, the special carinotaxy formula on the collum, coupled with certain details of gonopod structure (see also key below).

DESCRIPTION

Holotype with 43p+4a+ T, about 26 mm long and 1.6 mm wide; adult paratypes with 38p+4a+ T, about 22 mm long and 1.5 mm wide (♂), or with 43p+4a+ T, about 26 mm long and 1.6 mm wide (♀); with 53p+2a+ T, about 37 mm long and 2.0 mm wide (♀); with 56p+1a+ T, about 36 mm long and 2.2 mm wide (♀).

Coloration from uniformly pallid to marbled light yellow- or grey-brown, ozoporiferous cones often contrastingly dark brown; antennae, legs and venter yellowish to light yellow-brown; in adults, 8-12 blackish ocelli in a subtriangular eye patch on each side of head.

Antennae ( Fig. 26A View FIG ), gnathochilarium ( Fig. 26B View FIG ), postcollar constriction ( Fig. 25B View FIG ), dorsal crests and carinotaxy pattern on postcollar segments ( Fig. 25 View FIG ), segment’s texture and cross-section ( Fig. 25F View FIG ), paraprocts, ventral flaps of male segment 7, male legs 1-3 ( Fig. 26 View FIG C-E), penes ( Fig. 26D View FIG ) generally as in G. percostulifer n. sp., but collum ( Fig. 25A, B View FIG ) with a slightly deviating carinotaxy formula (I+2c+III-IV+5c +6a+pc+ma+pc+6a+5c+IV-III+2c+I); all tergal crests ( Fig. 25 View FIG A-E) on subsequent body segments higher, subdivided (2/2+I/i+3/3+I/i+2/2), ozoporiferous cones higher than wide; anterior half of paramedian dorsal crests particularly high, subtriangular on first 10-11 postcollar segments ( Fig. 25A, B View FIG ), thereafter somewhat lower ( Fig. 25 View FIG C-F); epiproct with a small but distinct tubercle at midway dorsally ( Fig. 25E View FIG ), very broadly rounded in caudal part in ventral view; hypoproct slightly but evidently emarginate mediocaudally; legs about as long as body diameter ( Fig. 25F View FIG ), always with a short accessory spine at base of claw ( Fig. 26 View FIG C-F).

Anterior gonopods ( Fig. 26G View FIG ) with coxosternum not plate-shaped but narrowed/elongate distally; telopodites straight. Posterior gonopods ( Fig. 26H, I View FIG ) also slightly elongate, flagella long and bare.

REMARKS

The juvenile paratype is haplopodous, with 32p+2a+T. As usual, the ozoporiferous cones on segments 5 and 6 are considerably larger than the others.

In the absence of clear troglomorphic traits, this species seems to be troglophilic at most. In addition, the cave is described as being “very small, full of bats and guano” (J. Lips in litt.).

Even though the gonopods of this new species are not very typical of the group concerned, their similarity to, and ready derivation from, the basic patterns is unquestionable. An assignment to Podoglyphiulus can be excluded on both morphological and zoogeographical grounds (see above).