Glyphiulus, GERVAIS, 1847 OF THE
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publication ID |
https://doi.org/ 10.5281/zenodo.5393630 |
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DOI |
https://doi.org/10.5281/zenodo.5469546 |
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persistent identifier |
https://treatment.plazi.org/id/B36487D8-FFE0-D51B-D31F-FF28FBCBFDEB |
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treatment provided by |
Marcus |
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scientific name |
Glyphiulus |
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KEY TO SPECIES OF GLYPHIULUS GERVAIS, 1847 OF THE View in CoL JAVANICUS- GROUP
1. At least some crests on collum more or less obliterated ( Figs 1A, B View FIG ; 3A, B View FIG ; 6A, B View FIG ; 10A, B View FIG ; 12A View FIG ) ...................................................................................................................... 2
— Crests on collum fully developed, no significant parts of collum smooth ( Figs 14A, B View FIG ; 17B View FIG ; 20A View FIG ; 22A, B View FIG ; 25A, B View FIG ) ........................................................................................ 8
2. Pleurosternal (= lowest) crests on segments 2-4 very prominent, arcuate ( Figs 1A View FIG ; 3A View FIG ); flagella of posterior gonopods missing; southern China.............................................. 3
— Pleurosternal crests on segments 2-4 not particularly prominent, as on subsequent segments ( Figs 6A View FIG ; 10A View FIG ); flagella of posterior gonopods present or absent ................................. 5
3. Body of adults 1.1-1.4 mm wide; carinotaxy formula of midbody segments 1/1+I/i+3+I/i+1/1 ( Fig. 1 View FIG A-E); male telopodites 1 slightly reduced, 4-segmented ( Fig. 2C View FIG ); cave in Yunnan Province, China.............................................................................. G. obliteratus View in CoL n. sp.
— Body of adults 1.6-2.2 mm wide; two divided crests below ozoporiferous cone; male telopodites 1 slightly reduced in size but 5-segmented ................................................ 4
4. Carinotaxy formula of midbody segments 2/2+I/i+3+I/i+2/2; body unpigmented, only ocelli black; cave in Guizhou Province, China.............................................. G. zorzini View in CoL
— Carinotaxy formula of midbody segments 2/2+I/i+3/3+I/i+2/2 ( Fig. 3A, B View FIG ); body and ocelli pigmented, at least partly dark; cave in Yunnan Province .... G. subobliteratus View in CoL n. sp.
5. Axial crest on midbody segments strongly reduced, carinotaxy formula 2/2+I/i+1/1+m+1/ 1+I/i+2/2 ( Fig. 6 View FIG ); legs slightly shorter than body diameter ( Fig. 6F View FIG ); anterior gonopod coxosternum with particularly high median outgrowths, devoid of a notch at distal margin ( Fig. 8G View FIG ); cave in North Vietnam........................................... G. mediobliteratus View in CoL n. sp.
— Axial crest on midbody segments not so strikingly reduced; legs subequal to or even longer than body diameter; anterior gonopod coxosternum deeply notched at distal margin; caves in Guizhou Province, China....................................................................................... 6
6. Carinotaxy formula of midbody segments 1/0+1+I/i+3/3+I/i+1+1/0 ( Fig. 12B, C View FIG ); male telopodites 1 normal, 5-segmented ( Fig. 13C View FIG ); anterior gonopods as in Figure 13G View FIG , posterior gonopods as in Figure 13H View FIG ......................................... G. parobliteratus View in CoL n. sp.
— Carinotaxy formula of midbody segments different; male telopodites 1 reduced in size but 5-segmented ( Fig. 11C View FIG ); anterior gonopods and posterior gonopods not as above ...... 7
7. Carinotaxy formula of midbody segments 1/0+1+I/i+3+I/i+1+1/0 ( Fig. 9 View FIG A-E); body and ocelli pigmented; medial outgrowth of anterior gonopod coxosternum folded ( Fig. 11G View FIG ) ................................................................................................... G. obliteratoides View in CoL n. sp.
— Carinotaxy formula of midbody segments 2/2+I/i+3/3+I/i+2/2; body and ocelli unpigmented medial outgrowth of anterior gonopod coxosternum not folded .... G. sinensis View in CoL n. comb.
8. Carinotaxy formula of collum typical for the granulatus View in CoL -group, I-VI+7a+pc+ma+pc+7a+VI-I, that of midbody segments 2+I/i+3/3+I/i+2 ( Fig. 14 View FIG A-E), texture of both lateralmost crests unusually micropunctate ( Fig. 15B View FIG ); male legs 1 and gonopods as in Figure 16C, G, H View FIG ; cave in Laos...................................................................................... G. costulifer View in CoL n. sp.
— Carinotaxy formula of collum not as above, both lateralmost crests usual, smooth; male legs 1 and gonopods not as above ............................................................................... 9
9. Carinotaxy formula of collum I-III+P+M+P+IV-I (i.e. all crests complete), that of midbody segments 2/2+I/i+3/3+I/i+2/2; anterior gonopod coxosternum without a deep notch at distal margin (unknown in G. formosus View in CoL ); male telopodites 1 strongly reduced, 1-3-segmented (unknown in G. formosus View in CoL ); flagella of posterior gonopods plumose distally (unknown in G. formosus View in CoL ) ........................................................................................ 10
— Carinotaxy formula of collum not as above, at least some of the crests being abbreviated; carinotaxy formula of midbody segments as above or different; anterior gonopod coxosternum often deeply notched at distal margin; male telopodites 1 usually less reduced; flagella of posterior gonopods, when present, bare or plumose ................................................. 13
10. Adult body about 1.5 mm wide; male femora 6 and 7 with a glandular adenostyle distoventrally; Zhejiang Province, China.................................................. G. recticullus View in CoL
— Body width at least 1.8 mm; male femora without adenostyles ................................. 11
11. Coloration light yellow-brown; only 12 ocelli in a subtriangular patch, black; adult body 1.8 mm wide; cave in Guangxi Province, China........................................... G. pulcher View in CoL
— Coloration darker, epigean species; fewer ocelli per eye patch; body about 2.5-2.6 mm wide ......................................................................................................................... 12
12. Nine ocelli in two rows per patch; Hong Kong (♀) .................................... G. formosus View in CoL
— Seven ocelli in three rows per patch; male telopodites reduced, 3-segmented; half-anterior gonopod with a subsecuriform coxosternum; Mt. Bana, Danang Province, Vietnam................................................................................................................... G. mediator View in CoL
13. Carinotaxy formula of collum I-V+P+ma+P+V-I, that of midbody segments 3/3+I/i+3+I/ i+3/3; not only male coxa 1 but also femur 1 with a median outgrowth; northern Thailand.................................................................................................................. G. siamensis View in CoL
— Carinotaxy formulae not as above; only male coxae 1 with a pair of paramedian processes ................................................................................................................................. 14
14. Carinotaxy formula of midbody segments 2+I/i+3+I/i+2 .......................................... 15
— Carinotaxy formula of midbody segments 2/2+I/i+3/3+I/i+2/2 ................................ 16
15. Carinotaxy formula of collum I-III+4c+5a+pc+ma+pc+5a+4c+III-I ( Fig. 22A, B View FIG ); male telopodites 1 reduced in size ( Fig. 24B View FIG ); half-anterior gonopod coxosternum narrowly notched at distal margin ( Fig. 24F View FIG ); cave in Guizhou Province, China................. ................................................................................................... G. paracostulifer View in CoL n. sp.
— Carinotaxy formula of collum I-IV+P+ma+P+IV-I; male telopodites 1 not reduced in size; half-anterior gonopod coxosternum broadly and barely emarginate at distal margin; cave in northern Vietnam...................................................................... G. vietnamicus View in CoL
16. Coloration entirely pallid, ocelli not visible; adult body up to about 1.0 mm wide; paramedian outgrowths of anterior gonopod half-coxosternum especially high and large (e.g., Fig. 21G View FIG ) ......................................................................................................... 17
— Coloration usually darker, ocelli always dark and discernible; body at least 1.3 mm wide; paramedian outgrowths of anterior gonopod half-coxosternum less conspicuous ...... 18
17. Ozoporiferous cones much wider than high; epiproct normal, smooth dorsally; male telopodites 1 reduced, 2-segmented; half-anterior gonopod with a particularly deep notch, such that both distal outgrowths are extremely prominent; posterior gonopods without flagella; Java, Indonesia (introduction from continental Southeast Asia?) ... G. javanicus View in CoL
— Ozoporiferous cones about as high as wide ( Fig. 20 View FIG A-E); epiproct unusual in showing two paramedian crests ( Fig. 20E View FIG ); male telopodites 1 normal, 5-segmented ( Fig. 21C View FIG ); half-anterior gonopod weakly emarginated at distal margin, only paramedian outgrowths prominent ( Fig. 21G View FIG ); posterior gonopods with flagella ( Fig. 21H View FIG ); cave in Laos.......... ..................................................................................................... G. percostulifer View in CoL n. sp.
18. Carinotaxy formula of collum particularly complex, I-II+3c+4a+5c+VI+7c+8a+pc+ma +pc+8a+7c+VI+5c+4a+3c+II-I ( Fig. 17B View FIG ); male telopodites 1 reduced in size, but still 5-segmented ( Fig. 19C View FIG ); half-anterior gonopod coxosternum subsecuriform ( Fig. 19G View FIG ); flagella of posterior gonopods plumose distally ( Fig. 19H View FIG ); cave in Laos........................ ..................................................................................................... G. subcostulifer View in CoL n. sp.
— Carinotaxy formula of collum I+2c+III-IV+5c+6a+pc+ma+pc+6a+5c+IV-III+2c+I ( Fig. 25A, B View FIG ); male telopodites 1 normal, 5-segmented ( Fig. 26C View FIG ); half-anterior gonopod coxosternum as in Figure 26G View FIG ; posterior gonopods as in Figure 26H View FIG , flagella bare; cave in Sichuan Province, China................................................................... G. intermedius View in CoL
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