Lesteva (Lestevina) sicula heeri Fauvel, 1871

Shavrin, Alexey V. & Zanetti, Adriano, 2021, Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula, Zootaxa 4966 (4), pp. 401-427 : 403

Lesteva (Lestevina) sicula heeri Fauvel, 1871

Lesteva heeri Fauvel, 1871a: 132 [= 1872: 106]; Portevin 1929: 438, Palm 1948: 119, Lohse 1956: 52

Lesteva sicula: Ganglbauer 1895: 714 , Reitter 1909: 184, Luze 1903: 185

Lesteva sicula heeri: Lohse 1960: 2, 1964: 59

Lesteva (Lestevina) heeri: Bordoni 1973: 234

Lesteva (Lestevina) sicula heeri: Tóth 1982: 98 , Zanetti 1987: 342, 2012: 69, Tronquet 2006: 33

Material examined. PORTUGAL: ALGARVE: 1 ♀: Serra Monchique E Monchique , 37°19’N 8°31’W. 530 m a.s.l. 10.04.2002. H. Meybohm leg. (cA); ESTREMADURA : 4 ♂♂, 1 ♀: Lisboa, Serra do Sintra. 24.05.1992. P. Wunderle leg. (cW) ; SPAIN: PAIS VASCO: 1 ♀: Irun. 10.02.2005. A. Anichtchenko leg. (cSh) ; 1 ♀: same data. 17- 19.03.2005 (cSh) ; 1 ♂, 3 ♀♀: same data. 09.04.2005 (cSh) ; 1 ♂, 2 ♀♀: same data. 05.2006 (cSh) ; 1 ♀: same data. 15.06.2006 (cSh); ARAGÓN : 2 ♂♂: Barbastro, Ariño de Cregenzán. 09.04.1995. D.A. Lott leg. ( BMNH) ; 1 ♀: Villanueva de Sigena. 09.04.1995. D.A. Lott leg. ( BMNH) ; 1 ♂: Guadalavier. 19.05.1994. D.A. Lott leg. ( BMNH) ; 1 ♂: Estrella ( BMNH); CASTILLA Y LÉON : 2 ♂♂: Sierra de Gredos , 10 km SSW Burgohondo, 40°21’15N 04°49’ 10W . 1250 m a.s.l. 25.03.2007. V. Assing leg. (cA); 2 ♂♂, 1 ♀: Jarandilla, Caseres. 16.06.1969. A. Comellini leg. ( MHNG) ; 1 ♂: Sierra de la Demanda. 2.8.1971. ( MHNG); EXTREMADURA : 2 ♂♂: Jarandilla. 29.06.1969. A. Comellini leg. ( MHNG); CASTILLA LA MANCHA : 1 ♂: Daimiel. 14.08.1969. A. Comellini leg. ( MHNG); CATALUñA : 1 ♂: Santa Perpètua de Gaiá. 08.04.1995. D.A. Lott leg. ( BMNH); VALENCIA : 1 ♂: El Toro. 950 m a.s.l. 07.04.1995. D.A. Lott leg. ( BMNH); ANDALUSÍA : 1 ♂: Sierra Magina , 37.72°N - 3.46°W. 2000 m a.s.l. 03- 04.05.2014. A.V. Shavrin leg. (cSch) ; 3 ♂♂, 2 ♀♀: Sierra Nevada, Capileira, Rio Nante , 36.99°N - 3.35°W. 1550 m a.s.l. 08.05.2014. A.V. Shavrin leg. (cSh) ; 3 ♂♂, 4 ♀♀: Capileira. 1600 m a.s.l. 22.03.1994. V. Assing leg. (cA) ; 1 ♀: Sierra Nevada , Puerto de la Ragua, 37.09°N - 3.02°W. 2000 m a.s.l. 08- 09.05.2014. A.V. Shavrin leg. (cSch) ; 3 ♂♂, 3 ♀♀: Sierra de Bermeja, Jubrique. 500 m a.s.l. 26.03.1994. V. Assing leg. (cA) ; 1 ♂, 1 ♀: Sierra Nevada, Guejar Sierra. 1200 m a.s.l. 28.09.1993. P. Wunderle leg. (cW) ; 1 ♀: San Pedro, Ronda. 14- 21.03.1990. Mattern leg. (cW) ; 4 ♂♂: Alpujarra Capileira. 1900 m a.s.l. 22.03.1994. P. Wunderle leg. (cW) ; 2 ♂♂, 1 ♀: Málaga, Jubrique. 500 m a.s.l. 26.03.1994. P. Wunderle leg. (cW) ; 1 ♂, 2 ♀♀: Sierra de Cazorla. 1400 m a.s.l. 06.10.1993. P. Wunderle leg. (cW) ; 1 ♂: Esteperona ( MHNG) ; 1 ♂, 1 ♀: Castille Puerto de Bejar. 800-1400 m a.s.l. 05-06.1957. G. Fagel leg. ( MHNG) ; 1 ♂: Puerto de la Ragua, Granada E. 1200 m a.s.l. 17.07.1971. A. Comellini leg. < Lesteva fontinalis infrasp. gustavi Det. Herman 2001> ( MHNG) .

Additional material. FRANCE: 1 ♂: ‘ Htes Pyrénées’ ( IRSN) ; MOROCCO: 1 ♂, 1 ♀: Grand Atlas Mts. , Oukaimeden env., 31°13’09.5’’N - 0.7°49’29.7’’W. 2200-2300 m a.s.l. 16- 17.06.2010. A. Anichtchenko leg. (cSh) ; 2 ♂♂: Alrene , 55 km S Marrakesh. 5000 ft. 26.iv.1961. P.N. Lawrence leg. ( BMNH) ; LOCALITY NOT SPECIFIED: 1 ♂: ‘ Fauvel’ , ‘ Pyrenäen’ ( SF) ; 1 ♂, 1 ♀: ‘ Pyren. ’, ‘ Fauvel’, ‘ heeri ’ ( SMNH) .

Remarks. Based on a study of material from several collections, L. sicula heeri was recorded by Lohse (1960) from the Pyrenees and central Spain. Horion (1963) recorded it from southern Spain (“Sierra Nevada, Kiesenwetter leg. Ca. 1850, 6 Ex. Z.S.M.”). Some studied specimens from southern Spain and Morocco have a paler body, welldeveloped hind wings and relatively wide elytra as in L. sicula sicula Erichson, 1840 , a subspecies distributed in Italy and Algeria. Lohse (1960) also noted this morphological similarity for specimens from Guadalupe, South Spain. This is probably morphological variability (from apterous to macropterous forms) of the species at its southern border of the distribution. The aedeagi of both subspecies have similar internal and external structures. A genetic analysis would be required for a clarification of the status and possible synonymy of both taxa.

The species is here recorded from Morocco for the first time.