Taxonomy of
Strandesia
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Bradleycypris obliqua ( Brady, 1868)
was placed in
Strandesia
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by Karanovic (2005), but we reject this decision, as characteristics of this species do not conform to those of
Strandesia
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s.s. As was mentioned above,
B. obliqua
, (the type species of this monospecific genus) has a strongly oblique carapace in frontal view and no inner list on the LV. Martens et al. (2008) accounted for 134 species in
Strandesia
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s.l. worldwide. Here, we retain a subset of these species in
Strandesia
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s.s. ( Table 5). Eleven species of
Strandesia
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are allocated to
Bradleystrandesia
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( Table 6) because of the presence of the Triebel’s loop in the dorsal branch. The following comments can be added.
Bradleystrandesia crassa ( Klie, 1939)
Bradleystrandesia dani ( George & Martens, 1993)
Bradleystrandesia decorata (Sars, 1903)
Bradleystrandesia lineata ( Victor & Fernando, 1981) Syn.
:
Strandesia amati ( Martens, 1984)
Bradleystrandesia parva ( Hartmann, 1964)
Bradleystrandesia tolimensis ( Roessler, 1990)
Bradleystrandesia trichurensis ( Victor et al., 1980)
Bradleystrandesia trispinosa ( Pinto & Purper, 1965) Syn.
:
Strandesia trispinosa galantis ( Broodbakker, 1983)
Bradleystrandesia tuberculata ( Hartmann, 1964)
Bradleystrandesia umbonata ( Victor & Fernando, 1981)
Bradleystrandesia weberi ( Moniez, 1892)
Strandesia amati ( Martens, 1984)
strongly resembles
Strandesia lineata Victor & Fernando, 1981
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, in valve ornamentation and in shape of the carapace. These species were considered by Martens (1984) as different taxa, owing to a number of different morphological features, e.g. the relative length of A2-claws, the length of the A2-aesthetasc, the shape of the Triebel’s loop and the number of rays on T1-respiratory plate. However, a comparison of the original descriptions of
S. amati
and
S. lineata
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, as well as new material from Thailand, reveals that these two taxa should be considered synonyms. Although there is some morphological variability in the chaetotaxy mentioned above, as well as in the setae on the T2-terminal segment, this is most likely owing to the fact that
S. lineata
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was described from last juvenile stage specimens ( Victor & Fernando 1981) in which limbs and setae are not yet completely developed.
Strandesia amati
is thus sunk in to the synonymy of
S. lineata
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and the latter species is transferred to
Bradleystrandesia
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.
Strandesia bicornuta
was described from India ( Hartmann 1964). The original description and illustrations were incomplete and even questionable in some features, so that the position of this enigmatic species remained uncertain. Martens & George (1992) redescribed the species on material from Kerala ( India). These authors mentioned that the species has more affinities with the South African
Cypricercus
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lineage than with
Strandesia
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, relying on the presence of anterior and posterior spines and the absence of a dorsal protuberance. However, after rechecking materials in the R.B.I.N.Sc. which were collected from Kerala State, India by Martens & George (1992) (specimes O.C. 1661–1670), it can be confirmed that this species belongs to
Strandesia
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s.s., as it has the typical characters of
Strandesia
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described above.
Strandesia trispinosa
, previously in
Cyprinotus
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, was described from southern Brazil by Pinto & Purper (1965). This species was divided into 2 subspecies,
S. trispinosa trispinosa
and
S. trispinosa galantis
, by Broodbakker (1983). Although the morphology of the soft parts of both forms was identical, Broodbakker (loc. cit) decided to erect this new subspecies because of the geographical distance. However, this view is ruled out since
S. trispinosa galantis
has recently also been recorded from the Paraná River ( Brazil) ( Higuti et al. 2007). Based on a comparison of the descriptions of these two subspecies and on a reexamination of material of
S. trispinosa
from Brazil, we conclude that the variation in length of the anterior and posterior spines is insufficient to warrant distinction between subspecies. Hence,
S. trispinosa galantis
is here considered a synonym of the nominotypical form. In addition, we lodge
S. trispinosa
in
Bradleystrandesia
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, because of the presence of the Triebel’s loop in the dorsal branch of the caudal ramus attachment.
Strandesia antetuberculata Hartmann & Petersen, 1985
Strandesia asymmetros Rome, 1962
Strandesia biwaensis Okubo, 2004
Strandesia bornemiszai Klie, 1935
Strandesia brteki Rybecki, 1988
Strandesia calapanensis Tressler, 1937
Strandesia canadensis ( Sars, 1926)
Strandesia caudata Klie, 1939
Strandesia cavernicola Broodbakker, 1983
Strandesia chondropherusa Rome, 1965
Strandesia clorocelis Anichini, 1967
Strandesia complexa Victor & Fernando, 1981
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Strandesia denticulata Tressler, 1950
Strandesia diversicolor Klie, 1938
Strandesia donnetii ( Baird, 1850)
Strandesia dorsoviridis McKenzie, 1966
Strandesia elatior ( Vavra, 1897)
Strandesia elliptica elliptica ( Sars, 1901)
Strandesia elliptica mayor Roessler, 1990
Strandesia elongata Hartmann, 1964
Strandesia feuerborni Klie, 1932
Strandesia flavescens Klie, 1932
Strandesia freyi Victor & Fernando, 1981
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Strandesia hystrix ( Furtos, 1933)
Strandesia intrepida Furtos, 1936
Strandesia itapeva Tressler, 1950
Strandesia kilimensis ( Daday, 1910)
Strandesia kraepelini ( Müller, 1906)
Strandesia laticauda ( Daday, 1910)
Strandesia mammarilorum mammarilorum Victor & Fernando, 1981
Strandesia mammarilorum sumatrana Victor & Fernando, 1981
Strandesia marmorata ( Brady, 1886)
Strandesia minuta Klie, 1936
Strandesia mulargiae Anichini, 1967
Strandesia mutica ( Sars, 1901)
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Strandesia obliqua Klie, 1940
Strandesia obtusata obtusata ( Sars, 1901) Syn.
:
Strandesia itapeva Tressler, 1950
Strandesia obtusata roessleri Martens & Behen, 1994
Strandesia ovalis Tressler, 1950
Strandesia pedroensis Tressler, 1950
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Strandesia phoenix De Deckker, 1981
Strandesia postica Rome, 1962
Strandesia prava Klie, 1935
Strandesia puncticulata ( Daday, 1910)
Strandesia purpurascens ( Brady, 1886)
Strandesia quasirotunda Hartmann & Petersen, 1985
Strandesia regularis Rome, 1962
Strandesia reticulata ( Daday, 1898)
Strandesia riograndensis Tressler, 1950
Strandesia rotunda ( Tressler, 1950)
Strandesia rouxi Mehes, 1939
Strandesia saetosa Hartmann, 1964
Strandesia santaeluciae ( Klie, 1935)
Strandesia sexpunctata Klie, 1932
Strandesia spinulosa Bronshtein, 1958
in Akatova, 1958
Strandesia striatoreticulata Klie, 1932
Strandesia sudanica Sywula, 1970
Strandesia taeniata ( Vavra, 1895)
Strandesia tenuicauda ( Brady, 1886)
Strandesia thermalis Rybecky, 1988
Strandesia tietensis Tressler, 1950
Strandesia towoetensis Tressler, 1937
Strandesia uenoi Klie, 1938
Strandesia ujijensis Rome, 1962
Strandesia unguiculata Brehm, 1938
Strandesia venezolana Broodbakker, 1983
Strandesia victori Harshey & Srinivasan, 1987
Strandesia vinciguerrae ( Masi, 1905) Syn.
:
Strandesia anterotundata Rome, 1977
Strandesia vittata (Sars, 1903)
Strandesia wierzejskii ( Grochmalicki, 1915)
Strandesia wolterecki Tressler, 1937
The exact position of most of the other
Strandesia
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and
Cypricercus
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species remains unclear, mainly because of the incomplete descriptions of these species. In most cases, no account is provided of the appearance of the α-seta on the Md-palp and of the d-seta on the T1, which are significant taxonomic features of this group. These uncertain species are listed in Tables 4 and 7 for
Cypricercus
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and
Strandesia
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, respectively.