Soyedina alexandria Grubbs, 2006

Grubbs, Scott A. & Baumann, Richard W., 2019, Soyedina Ricker, 1952 (Plecoptera: Nemouridae) in the eastern Nearctic: review of species concepts, proposed morphology-based species groups, and description of a new species from North Carolina, Zootaxa 4658 (2), pp. 223-250 : 225-227

Soyedina alexandria Grubbs, 2006

Cumberland Forestfly

http://lsid.speciesfile.org/urn:lsid: Plecoptera .speciesfile.org:TaxonName:605 8

(Figs. 1–8, 79)

Soyedina alexandria Grubbs 2006:43 . Holotype ♂ (INHS), spring into Pinewood Branch , Leipers Fork , West Harpeth River, 4 km W Leipers Fork, Williamson Co., Tennessee, USA

Distribution. USA: AL, MS, TN ( DeWalt et al. 2019).

Material examined. USA, Alabama, Limestone Co., tributary to East Fork Scarce Grease Branch, Sugar Creek , Smith Hollow , 2 km SE Lester, 34.99087, -87.12323, 18 February 2008, S.A. Grubbs, 5 males, 1 female ( WKUC) ; same but 34.98164, -87.13036, 18 February 2008, S.A. Grubbs, 1 male, 8 females ( WKUC) ; Jackson Co., spring into Larkin Fork, Rte. 65, 3 km SE Francisco , 34.97997, -86.23382, 18 February 2006, S.A. Grubbs, 2 males, 1 female ( WKUC) ; same but 19 February 2007, S.A. Grubbs, 1 male ( WKUC) ; spring into unnamed tributary to Big Coon Creek, 4 km NE Hytop, Skyline Wildlife Management Area , 34.94133, -86.06400, 19 February 2007, S.A. Grubbs, 4 males, 2 females ( WKUC) . Mississippi, Tishomingo Co., tributary Sandy Creek , CR 355, 6 February 2010, A. Harrison, L. Little, 2 males ( BPSC) . Tennessee, Cheatam Co., tributary to Harpeth River, Rte. 49, 6 km W Ashland City , 36.27551¸-87.12691, 1 March 2003, S.A. Grubbs, 2 males, 5 females (paratypes, WKUC) ; Sumner Co., spring into Liggett Branch, Station Camp Creek , 6 km SE White House, 36.43708¸-86.59967, 1 March 2003, S.A. Grubbs, 3 males, 8 females (paratypes, WKUC) ; Williamson Co., spring into Pinewood Branch, Leipers Fork , Rte. 46, 4 km W Leipers Fork , 35.89440, -87.03878, 16 March 2003, S.A. Grubbs, 1 male, 4 females (paratypes, WKUC) ; same but 5 March 2007, S.A. Grubbs, 1 male ( WKUC) .

Remarks. Adults were described and illustrated in Grubbs (2006); the larva remains undescribed. The combination of the asymmetric dorsal (Figs. 1–2, 4, 6) and ventral sclerites (Figs. 1–2), plus the inner epiproct member that is flanged and directed laterally with the flange lip enlarged apically (Figs. 2–5), aligns this species closely with S. calcarea and S. vallicularia .

The known distribution of S. alexandria is along a narrow latitudinal and longitudinal band from Cumberland Plateau region in northwestern Alabama westward to northern Mississippi and northward through middle Tennessee ( Fig. 79 ).

Figures

FIGURE 79. Distribution maps of eastern Nearctic Soyedina based on material examined during this study plus valid literature data. 79, S. alexandria, S. calcarea, and S. vallicularia.