Neoperla (Borneella), 2023

Zwick, Peter, 2023, A new classification of genus Neoperla and systematic studies of other Perlinae (Plecoptera: Perlidae), Zootaxa 5339 (2), pp. 101-131 : 111

2.4.2.1. Neoperla (Borneella) , new subgenus

Type species: N. bilobata Zwick, 1986 ; additional included species: N. borneensis (Enderlein, 1909) , N. furcata Zwick, 1986 , N. furcifera Klapálek,1909 , N. harina Navás, 1929 , N. naviculata Klapálek,1909 .

Male: T7 with a deeply divided flat process or a pair of adjacent processes, sharp spines (modified sensilla basiconica?) along the margins. T8 flat with a band of spinules under each projection of T7, small spinule groups or pilose humps on T9. HT10 with a caudal extension that bears grouped setulae along the medial edge (e.g., Figs. 46, 48 ). Penis base membranous and soft except for a small ventral and an elongate pointed dorsal sclerite. The endophallus varies from short and curved with some protruding lobes ( Fig. 49 ), to very large bizarre shapes with several large lobes or long fingers, the largest spines at the tips.

Known females: No or a short subgenital plate. Vagina with minute pale spicules in center, otherwise unmodified, topped by a stack of approximately circular folds. The short spermathecal duct and the sausage-shaped spermatheca insert on top.

Known eggs are not striate but have irregular punctures or polygones, with or without collar, anchors are multifilamentous, each filament ends in a small adhesive body ( Fig. 47 ).

Figures

FIGURES 46–49. Neoperla (Borneella) spp.: 46–47, N. (B.) naviculata Klapálek, 1909, male abdominal tip and egg with multifilamentous anchor; 48–49, N. (B.) bilobata Zwick, 1986, male abdominal tip and penis with everted endophallus; not to scale. From Zwick (1986).