Phytocrene anomala Merr., 1840

Rio, Cédric Del, Stull, Gregory W. & Franceschi, Dario De, 2020, Survey of the fruits and endocarps of Icacinaceae (Lamiids, Icacinales), European Journal of Taxonomy 645, pp. 1-130 : 62

Phytocrene anomala Merr.

Fig. 20.19–20.26

Material examined

Specimen used for endocarp and fruit description

BORNEO • 1920; M. Ramos 1840; L [ L.2289425 ].

MALAYSIA • 18 Jun. 1986; Patrick et al. SAN 118639 ; K [ K000183648 ].

Description

FRUIT. Elliptical, slightly asymmetrical and accrescent at the apex, yellow-orange when mature. Epicarp with clusters of yellow hairs. Calyx persistent. Length 20–25 mm, width 11–15 mm, thickness 8.0– 10.1 mm.

ENDOCARP. Cream, elliptical in lateral view, lenticular in transverse section, length ca 14.4 mm, width ca 10 mm, thickness ca 6 mm. Marginal keel encircling the endocarp in the plane of symmetry, lacking except for a thin trace observable in the upper part. Apex slightly asymmetrical and acute in lateral view; base rounded, symmetrical. Outer surface of the endocarp pitted. Pits circular to elongate (concave area), ca 0.8–1.7 mm in diameter, representing most of the surface of the endocarp, more or less organized longitudinally with ca 8–10 pits longitudinally and 6–7 pits transversally (ca 35–45 pits per face). Each pit with a central dimple or one freely ending ridge inside. Pits without tubercles. Endocarp wall ca 330 µm thick excluding pits (114–154 µm thick including pits). Endocarp wall (excluding pits) with three cell layers: outermost layer with 3–5 rows of anticlinally oriented cells, cells 12.3–24.5 µm in length; middle layer with 9–10 rows of periclinally oriented cells, cells 10.2–14.2 µm in width; innermost layer with a single row of periclinally oriented cells, cells 13.0– 20.7 µm in width, lining the locule surface with inflated cells. Locule with slightly rounded protuberances due to the thickening of the wall under the pits.