Iodes philippinensis Merr.

Rio, Cédric Del, Stull, Gregory W. & Franceschi, Dario De, 2020, Survey of the fruits and endocarps of Icacinaceae (Lamiids, Icacinales), European Journal of Taxonomy 645, pp. 1-130 : 42-44

Iodes philippinensis Merr.

Fig. 15.1–15.9

Material examined

Specimen used for endocarp and fruit description

PHILLIPINES • 1916; A.D.E. Elmer 16418; P [ MNHN-P-P04504850 ].

Description

FRUIT. Oblong, red when mature. Epicarp puberulent-strigose, with yellow simple hairs with granular ornamentation and uncinate hairs, shriveled when dry, revealing the underlying endocarp ridges. Mesocarp ca 50 µm thick when dry. Length 11–15 mm, width 7.5–12.0 mm, thickness 5–10 mm.

ENDOCARP. Brown, oblong in lateral view, hexagonal in transverse section, length ca 12.4 mm, width ca 7.6 mm, thickness ca 6.5 mm. Keel surrounding the endocarp in the plane of symmetry, with the thicker margin containing the primary vascular bundle embedded within the endocarp wall. Apex asymmetrical and almost flat with two uncinate protuberances in lateral view; base flattened and cleft on the opposite side. Outer surface of the endocarp ridged with two types of ridges. Primary ridges large and rectangular, composed of two main longitudinal ridges, running from the subbasal part up to the subapical part of the endocarp length and forming a diffuse pattern with only 2–3 lateral freely ending ridgelets. Secondary ridges thin and rounded, delimiting a dense reticulum enclosing numerous areoles. Endocarp possessing a symmetrical pair of horn-like protrusions positioned on the bigger uncinate protuberance at the apex. Vasculature of the endocarp resting on ridges. Endocarp wall 229–282 µm thick excluding ridges (285– 296 µm including secondary ridges and 360–396 µm including primary ridges). Endocarp wall with four cell layers: outermost layer with ca 5 rows of isodiametric cells, cells 12.5–16.5 µm in diameter, followed by a layer with ca 6 rows of isodiametric to anticlinally oriented cells, cells 21.4–28.3 µm in width, then a layer with ca 5 rows of periclinally oriented cells, cells 8.1–11.9 µm in width; innermost layer with one row of periclinally oriented cells, cells 6.7–6.9 µm in width, covering the surface of the locule (detail of the surface unknown). Locule lacunate.