Iodes ovalis Blume

Rio, Cédric Del, Stull, Gregory W. & Franceschi, Dario De, 2020, Survey of the fruits and endocarps of Icacinaceae (Lamiids, Icacinales), European Journal of Taxonomy 645, pp. 1-130 : 39-40

Iodes ovalis Blume

Fig. 14.10–14.18

Material examined

Specimen used for endocarp and fruit description

VIETNAM • Hiep HLF 203; MO [ MO-2080935 ].

Other material

INDONESIA • “ Celebes ” [Sulawesi]; 1863; Kunz ?; P [MNHN-P-P04504863] .

Description

FRUIT. Elliptical, cleft at the base, red when mature. Epicarp strigose with yellow simple hairs with granular ornamentation and small papillae-like hairs lining the fruit, shriveled when dry, revealing the underlying reticulum of endocarp ridges. Mesocarp 142–242 µm thick when dry. Length 15–25 mm, width 10.0– 11.4 mm, thickness 8–9 mm.

ENDOCARP. Cream, elliptical in lateral view, lenticular in transverse section, length ca 17.8 mm, width ca 11 mm, thickness ca 8.6 mm. Keel surrounding the endocarp in the plane of symmetry, with the thicker margin containing a vascular bundle embedded within the endocarp wall. Apex asymmetrical and acute in lateral view; base flattened and cleft. Outer surface of the endocarp ridged and faintly rugose, with 3–4 main longitudinal ridges delimiting a reticulate pattern, enclosing 3–4 large polygonal areoles. Ridges thin, rectangular, with a median channel, interconnected with some very short freely ending ridgelets. Rugosities distributed over the entire surface. Endocarp possessing a symmetrical pair of horn-like protrusions positioned eccentrically and subapically on the endocarp faces, each with a central pit. Endocarp wall 314–375 µm thick excluding ridges (ca 490 µm thick including ridges). Endocarp wall with three cell layers: outermost layer with 16–19 rows of isodiametric, occasionally anticlinally oriented cells, followed by a layer with 6–11 rows of periclinally oriented cells; innermost layer with one row of periclinally oriented cells, lining the locule surface with regularly spaced and rounded papillae; papillae 9.2–11.8 µm (av. 10.1 µm) in diameter, at least 220 papillae per 0.25 mm 2. Locule lacunate.