Rhyticaryum Becc.

Rio, Cédric Del, Stull, Gregory W. & Franceschi, Dario De, 2020, Survey of the fruits and endocarps of Icacinaceae (Lamiids, Icacinales), European Journal of Taxonomy 645, pp. 1-130 : 98

Genus Rhyticaryum Becc.

Figs 31.10–31.25, 32, 33.1–33.18

Description

FRUIT. Elliptical, asymmetrical at the apex, laterally compressed, red when mature. Epicarp strigose, with simple hairs with granular ornamentation or glabrous, shriveled when dry, revealing the underlying reticulum of endocarp ridges. Calyx persistent.

ENDOCARP. Brown-cream, elliptical in lateral view, lenticular in transverse section. Keel surrounding the endocarp in the plane of symmetry. Apex asymmetrical in lateral view, with pair of subapical pores, occasionally absent; base symmetrical. Outer surface of the endocarp ridged, rarely rugose; ridges diffuse or delimiting a reticulate pattern enclosing polygonal areoles with freely ending ridgelets. Ridges rounded, occasionally sharp. Vasculatures of the endocarp resting on the ridges. Endocarp primary vascular bundle outside the endocarp wall or in a channel on the keel. Endocarp wall at least with one layer with rows of periclinally oriented cells and another with rows of anticlinally oriented cells. Locule surface covered with large to rounded and regularly spaced papillae. Locule lacunate.