Rhyticaryum oleraceum Becc.

Rio, Cédric Del, Stull, Gregory W. & Franceschi, Dario De, 2020, Survey of the fruits and endocarps of Icacinaceae (Lamiids, Icacinales), European Journal of Taxonomy 645, pp. 1-130 : 102-104

Rhyticaryum oleraceum Becc.

Fig. 33.1–33.9

Material examined

Specimen used for endocarp and fruit description

INDONESIA • Moluccas isl.; 1938; P. Buwalda 4471; L [ L.2293162 ].

Description

FRUIT. Epicarp glabrous; mesocarp 260–270 µm thick when dry. Length 17–26 mm, width 10.0– 15.7 mm, thickness 7.4–10.0 mm.

ENDOCARP. Cream, length ca 23.6 mm, width ca 15.7 mm, thickness ca 8.7 mm. Keel surrounding the endocarp in the plane of symmetry. Apex slightly inflated and asymmetrical in lateral view; base rounded, symmetrical. Outer surface of the endocarp ridged with three main ridges longitudinally. Ridges rounded and thin, forming a diffuse pattern, with two ridges spanning from the apex to nearly the base and a median ridge spanning from the base to nearly the apex. Primary vascular bundle positioned outside the endocarp wall. Endocarp wall 157–188 µm thick excluding ridges (ca. 420 µm including ridges). Endocarp wall with three cell layers: outermost layer with 5–7 rows of anticlinally oriented cells, cells 17.2–29.4 µm in length, followed by a layer with 2–3 rows of periclinally oriented cells, cells 15.4–24.0 µm in width; innermost layer with one row of periclinally oriented cells, cells 4.3–6.0 µm in width, lining the locule surface with regularly spaced and large papillae; papillae 8.9–28.3 µm (av. 15.0 µm) in diameter.