Drassodella melana Tucker, 1923

Mbo, Zingisile & Haddad, Charles R., 2019, A revision of the endemic South African long-jawed ground spider genus Drassodella Hewitt, 1916 (Araneae: Gallieniellidae), Zootaxa 4582 (1), pp. 1-62: 25-28

publication ID

https://doi.org/10.11646/zootaxa.4582.1.1

publication LSID

lsid:zoobank.org:pub:9DC61794-4BD7-4F6D-BB8C-84D9855C8151

persistent identifier

http://treatment.plazi.org/id/B74F87AB-9E79-403A-53B3-CFF867D39F6F

treatment provided by

Plazi

scientific name

Drassodella melana Tucker, 1923
status

 

Drassodella melana Tucker, 1923  

Figs 64 View FIGURES 60–77 , 80 View FIGURES 78–92 , 99, 100 View FIGURES 93–110 , 126–130 View FIGURES 126–130

Drassodella melana Tucker, 1923: 310   , fig. 38 (Holotype ♀: SOUTH AFRICA: KwaZulu-Natal Province: Mfongosi, 28°42'S, 30°50'E, II.1918, leg. W.E Jones, SAM ENW-X008521—examined).

Diagnosis. Drassodella melana   females are most similar to those of D. venda   sp. nov., but can be distinguished by the anterior hood having only a slightly convex anterior margin ( Fig. 64 View FIGURES 60–77 ), while strongly convex in D. venda   sp. nov. ( Fig. 69 View FIGURES 60–77 ), and by lateral margins of ST I being slightly broader than those of ST II, while ST II are slightly broader in D. venda   sp. nov.. Males of D. melana   are most similar to those of D. guttata   sp. nov., but can be distinguished by the absence of abdominal markings (white spots present in D. guttata   sp. nov.), the spatulate rather than subtriangular shape of the ventral prong in prolateral view (compare Fig. 128 View FIGURES 126–130 with 118), and the RTA that in retrolateral view is more slender with a slightly curved tip as opposed to broad basally and tapering to a point distally (compare Fig. 130 View FIGURES 126–130 with 120).

Description. Female (Ngome Nature Reserve, NCA 94/364). Measurements: CL 2.52, CW 1.74, AL 3.56, AW 2.1, TL 6.24 (4.90–9.85), FL 0.41, SL 1.37, SW 1.12, CH 0.13, AME-AME 0.08, AME-ALE 0.16, ALE-ALE 0.21, PME-PME 0.10, PME-PLE 0.10, PLE-PLE 0.44, PERW 0.52, MOQAW 0.20, MOQPW 0.21, MOQL 0.28.

Length of leg segments: I 1.52 + 0.76 + 1.12 + 1.06 + 0.86 = 5.32, II 1.32 + 0.64 + 0.98 + 0.96 + 0.82 = 4.72, III 1.20 + 0.56 + 0.84 + 1.14 + 0.80 = 4.54, IV 1.80 + 0.80 + 1.42 + 1.94 + 0.96 = 6.92.

General appearance as in Fig. 99 View FIGURES 93–110 . Carapace dark brown, with black mottling; thin white line of feathery setae diverging from fovea towards fourth legs, not reaching margin; few white feathery setae from eye region to posterior fovea. Clypeus height equal to 1.63 AME diameter; AME smaller than ALE; AME separated by distance equal to their diameter, AME separated from ALE by distance twice AME diameter; PME smaller than PLE; PME separated by distance equal to 1.43 their diameter, PME separated from PLE by distance equal to 1.43 PME diameter. Chelicerae dark-brown, orange around fang; promargin with three teeth, median tooth largest; retromargin with two teeth, distal tooth largest. Endites with two light patches anteriorly and posteriorly, dark fringe mesally, cream around maxillar hair tuft. Labium as broad as long. Sternum with elongate setae evenly spread across surface. Legs dark-brown, with mottling proximally, distal segments orange. Leg spination: femora: I do 2 pl 1, II do 2 pl 1, III do 6, IV do 2; patellae: I spineless, II rl 1 vt 1, III vt 1, IV rl 1; tibiae: I & II spineless, III pl 2 do 2 rl 2 plv 2 rlv 2 vt 2, IV pl 2 do 2 rl 2 plv 2 rlv 2 vt 1; metatarsi: I & II spineless, III pl 2 rl 2 plv 2 rlv 2 vt 3, IV pl 2 do 1 rl 4 plv 2 rlv 3 vt 3; palpal spination: femora pl 2; patellae do 1; tibiae pl 1 do 2 plv 2; tarsi do 1. Abdomen uniformly dark brown ( Fig. 99 View FIGURES 93–110 ); two lateral sclerites above booklungs; venter light brown. Spinnerets light brown. Epigyne with deep anterior hood, anterior and posterior margins convex, lateral ends terminating abruptly; lateral hoods curved, close to copulatory openings; ST II orange, connecting ducts and ST I orangebrown; ST II slightly larger than ST I, ST II elongate-oval ( Figs 64 View FIGURES 60–77 , 126, 127 View FIGURES 126–130 ).

Male (Ngome State Forest, NCA 2002/420). Measurements: CL 2.10, CW 1.58, AL 2.32, AW 1.16, TL 4.72 (4.20–6.90), FL 0.39, SL 1.12, SW 0.98, CH 0.13, AME-AME 0.07, AME-ALE 0.02, ALE-ALE 0.19, PME-PME 0.07, PME-PLE 0.10, PLE-PLE 0.37, PERW 0.44, MOQAW 0.15, MOQPW 0.19, MOQL 0.24.

Length of leg segments: I 1.52 + 0.76 + 1.12 + 1.06 + 0.86 = 5.32, II 1.32 + 0.64 + 0.98 + 0.96 + 0.82 = 4.72, III 1.20 + 0.56 + 0.84 + 1.14 + 0.80 = 4.54, IV 1.80 + 0.80 + 1.42 + 1.94 + 0.96 = 6.92.

General appearance as in Fig. 100 View FIGURES 93–110 . Carapace dark-brown with black mottling; white feathery setae spread across surface, forming distinct medial line that diverges behind fovea, forming upsilon shape; male with more dense white feathery setae than female. Clypeus height equal to 2.17 AME diameter; AME smaller than ALE; AME separated by distance slightly larger than their diameter, AME separated from ALE by distance 0.33 AME diameter; PME equal to PLE; PME separated by distance equal to 1.17 their diameter, PME separated from PLE by distance equal to 1.67 PME diameter. Chelicerae dark-brown; promargin with three evenly spaced subequal teeth; retromargin with two widely separated subequal teeth; base of fang orange. Endites and labium as for female. Sternum with dark-brown markings; setae spread, strongly concentrated around edges. Legs with dark-brown proximal segments, distally segments orange; legs with white feathery setae on proximal segments only. Leg spination: femora: I do 2 rl 1, II do 3, III do 5 rl 1, IV do 4; patellae: spineless; tibiae I & II spineless, III pl 2 do 2 rl 2 plv 2 rlv 2 vt 2, IV pl 2 do 2 rl 2 plv 2 rlv 2 vt 2; metatarsi: I & II spineless, III do 2 rl 3 rlv 2 plv 2 vt 2, IV do 1 pl 2 plv 2 vt 3 rl 4 rlv 3; palpal spination: femora do 3; patellae pl 3 do 1 rl 1; tibiae do 3 rl 2 plv 2; tarsi do 1. Abdomen brown ( Fig. 100 View FIGURES 93–110 ); sub-oval sclerite laterally above booklungs; venter light brown; two rows of tiny sclerites running from epigastric furrow to spinnerets. Spinnerets pale. Palp yellow-brown; RTA moderately long, largely obscured in ventral view by retrolateral heel of cymbium; in retrolateral view triangular, with broad base and slightly curved tip; cymbium quite narrow, 1.8 times longer than wide, narrowed to rounded point distally; ventral embolic prong spatulate; dorsal prong narrow, slightly curved in prolateral view, aligned with palpal axis in ventral view; median apophysis narrow, hook-shaped ( Figs 80 View FIGURES 78–92 , 128–130 View FIGURES 126–130 ).

Additional material examined. SOUTH AFRICA: KwaZulu-Natal Province: 15 km ENE of Louwsberg on R69, 27°31.801'S, 31°25.386'E, 1025 m a.s.l., 5. II.2005, C. Haddad (leaf litter, Eucalyptus   plantation), 1♂ 2♀ (NCA 2016/2822); Good Hope, Midlands, 28°15'S, 30°42'E, I.2014, leg. I. Yekwayo, 1♀ (2015/1550); Ithala Game Reserve, Ntshodwe Camp, 27°32.699'S, 31°16.911'E, 26.I.2014, leg. C. Haddad & Z. Mbo (sifting leaf litter, short forest), 2♀ ( NCA 2013/5006); Same as previous but 27.I.2014, 3♀ ( NCA 2013/4938); Ithala Game Reserve, Onverdacht picnic site, 27°31.967'S, 31°18.984'E, 29.I.2014, leg. C. Haddad (base of grass tussocks), 1♀ ( NCA 2013/5094); Jackson’s Falls on Mhlatuzana River, 29°48'S, 30°45'E, 18. XII.1990, leg. V. & B. Roth, 1♀ (CAS, CASENT 9072379); Krantzkloof Nature Reserve, 29°46'S, 30°49'E, 24. VII.2002, leg. C. Haddad (leaf litter on hillside), 1♂ 1♀ (NCA 2002/420); Krantzkop, 28°58'S, 30°52'E, XI.1917, leg. K.H. Barnard, 3♀ (SAMC ENW- B003610); Vryheid, Ngome State Forest, 27°49'S, 31°26'E, I.1992, leg. M. van der Merwe (pitfall traps, grass), 2♂ 6♀ (NCA 94/364); Same data as previous but II.1992, 2 ♂ 3♀ (NCA 94/366), 2♀ 1imm. (NCA 94/369), 2♀ (NCA 94/372); Same data as previous but III.1992, 1 ♀ (NCA 94/394); Same data as previous but IV.1992, 1 ♀ (NCA 94/ 371), 3♂ 8♀ (NCA 94/380); Same data as previous but V.1992, 2 ♂ 1 imm. (NCA 94/378); Same data as previous but VI.1992, 1 ♂ 1 imm. (NCA 94/368), 1♀ (NCA 94/391); Same data as previous but VII.1992, 1 ♂ 1♀ 2 imm. (NCA 94/363), 3♂ 3♀ (NCA 94/390); Same data as previous but VIII.1992, 2 ♂ 2♀ (NCA 94/377), 1♂ (NCA 94/ 385); Same data as previous but IX.1992, 1 ♂ 8♀ (NCA 94/367), 1♂ (NCA 94/384), 1♂ 1♀ (NCA 94/388); Same data as previous but X.1992, 1 ♂ 2♀ (NCA 94/374), 1♂ 4♀ 2 imm. (NCA 94/392); Same data as previous but XI.1992, 7 ♀ (NCA 94/381); Same data as previous but XII.1992, 3 ♂ 8♀ (NCA 94/382); Same data as previous but I.1993, 5 ♂ 8♀ (NCA 94/383), 3♀ (NCA 94/365), 5♂ 8♀ (NCA 94/3813), 5♂ 9♀ (NCA 94/386); Vryheid, Ngome State Forest, 27°49'S, 31°26'E, IX.1992, leg. M. van der Merwe (pitfall traps, dense forest), 1♀ (NCA 94/422), 1♂ (NCA 94/507); Vryheid, Ngome State Forest, 27°49'S, 31°26'E, IX.1992, leg. M. van der Merwe (pitfall traps, pine plantation), 1♀ (NCA 94/379); Same data as previous but I.1993, 1 ♂ (NCA 94/370). Limpopo Province: Lekgalameetse Nature Reserve, The Downs, 24°08'S, 30°18'E, 19.I.2009, leg. S.H. Foord, 1♂ (NCA 2015/3708). Mpumalanga Province: Mariepskop State Forest, Drakensberg, 23°35.06'S, 30°51.81'E, 1350 m a.s.l., 13.X.1999, leg. D. Ubick & S. Prinsloo, 1♀ (CAS, CASENT 9072381); Same locality, XI.2013, leg. L. Taylor, 2♂ 1♀ 1 imm. (NCA 2015/4009), 1♀ (NCA 2015/4915), 2♂ 3♀ (NCA 2015/4918), 2♂ 3♀ (NCA 2015/4974); Same data as previous but I.2013, 3 ♀ 2 imm. (NCA 2015/4917); Same data as previous but II.2014, 1 ♀ (NCA 2015/4916).

Distribution. Found in the KwaZulu-Natal, Mpumalanga and Limpopo Provinces of South Africa ( Fig. 140 View FIGURE 140 ).

Kingdom

Animalia

Phylum

Arthropoda

Class

Arachnida

Order

Araneae

Family

Gallieniellidae

Genus

Drassodella

Loc

Drassodella melana Tucker, 1923

Mbo, Zingisile & Haddad, Charles R. 2019
2019
Loc

Drassodella melana

Tucker, R. W. E. 1923: 310
1923