Bessierus doloris Thomas & Orth, 2000 in Thomas, Orth, Horeau & Dominique (2000)

Cruz, Paulo Vilela, Gonçalves, Inês C., Mariano, Rodolfo & Hamada, Neusa, 2020, One less monotypic genus in Leptophlebiidae (Ephemeroptera): A new species of Bessierus Thomas & Orth and new record from Brazil, Zootaxa 4820 (1), pp. 195-200 : 196

Bessierus doloris Thomas & Orth, 2000 in Thomas, Orth, Horeau & Dominique (2000)

( Figure 2 )

Bessierus doloris Thomas & Orth 2000: 50 , (nymph); Domínguez et al., 2019: 379–381, figs. 2–3 (male imago); Neto et al. 2020: 345.

Material examined. 1 male subimago; Amapá; M. Amapá; Rio Amapá Grande, Cachoeira Grande ; 3 m alt.; 02º09’43.1”N / 50º55’17.3”W; 07.viii.2011; A. Pes, P. V . Cruz, A. Fernandes, N. Hamada cols; light trap; INPA .

Diagnosis. Male imago: (1) penis lobes widely separated from each other ( Figure 2c ); (2) base of each penis lobe, on outer margin, without expansion; (3) telopenis broad; (4) inner margin of basal 1/3 of forceps segment I smoothly rounded.

Figures

FIGURE 2. Male subimago of Bessierus doloris. a. forewing; b. body, dorsal view; c. genitalia, ventral view.