Goeldia goytacazes, Almeida-Silva & Brescovit, 2024

Almeida-Silva, Lina M. & Brescovit, Antonio D., 2024, Unraveling the mysteries of Goeldia Keyserling, 1891: revision, description of seven new species and first record from USA (Araneae: Titanoecidae), Zootaxa 5428 (2), pp. 151-193 : 168-170

Goeldia goytacazes sp. nov.

( Figs 12 ; 27 )

Type material. Holotype male, paratype female from Campos de Goytacazes (21°45’14”S 41°19’26”W), Rio de Janeiro, Brazil, 24.IX.2003, C.C.L. Teixeira coll., deposited at MCN 39868 ; Paratype female with same data as holotype, deposited at MCN 43434.

Additional material examined. None.

Etymology. The specific name is a noun in apposition in honor to the indigenous people of the Goytacazes tribe, who lived in the type locality.

Diagnosis. Goeldia goytacazes resembles G. mirim by the general shape of the male palp and epigynum ( Fig. 16 ), but differs by the patellar apophysis pointing posteriorly ( Fig. 12C ); by the enlarged base of the MLT ( Fig. 12C ) and tegular process lanceolate ( Fig. 12A ) in the male palp. The female epigynum is also similar but in G. goytacazes it covers almost completely the copulatory duct wall ( Fig. 12E ) and the vertical spermathecae is much shorter than the copulatory duct ( Fig. 12F ).

Description. Male (Holotype). Coloration: carapace dark brown. Sternum and labium black, endites brown. Leg coxae and femora black, other leg segments yellowish brown except distal half of patella and tibia III and IV black and metatarsus IV grayish. Opisthosoma grayish brown. Total length 3.15. Carapace: lenght 1.59; width 1.16; height 0.75. Opisthosoma: length 1.65; width 1.38. Clypeus: height 0.13. Eye diameters and interdistances: AME 0.04; ALE 0.08; PME 0.06; PLE 0.06; AME–AME 0.08; AME–ALE 0.07; PME–PME 0.1; PME–PLE 0.15; AME–PME 0.08; ALE–PLE 0.07. Chelicerae paturon: length 0.65. Leg measurements: I: femur 1.23/ patella 0.50/ tibia 1.05/ metatarsus 0.98/ tarsus 0.62/ total 4.37; II: 1.13/ 0.48/ 0.83/1.03/ 0.48/ 3.93; III: 1.05/ 0.4/ 0.93/ 0.75/ 0.34/ 3.46; IV: 1.25/ 1.20/ 0.5/ 1.06/ 0.44/ 4.45. Leg formula 4123. Leg spination: femur I pap; tibia I v1r-2(1r- 1m)- 2(1r- 1m)-0, p0-1-0, r0-1-0-1-0; III–IV: p1ap, r1ap; metatarsus I v1r-1r- 1m, p1ap, r1ap; II v1r-0-1r- 1m, p1ap, r1ap; IV v1r-1p-1map, p0-1-1ap, r1ap. Patellar apophysis pointing dorsally and posteriorly ( Fig. 12C–D ). RLT with an extra basal fold ( Fig. 12C ). Tegular process as in illustration ( Fig. 12A ). Pars pendula thickening subdistal ( Fig. 12D ).

Female (Paratype): Coloration as in male except, femora black, tarsus I and II reddish brown and other leg articles yellowish brown. Total length 3.84. Carapace: length 1.80; width 1.24; height 0.75. Opisthosoma: lentgth 1.24; width 1.48. Clypeus: height 1.40. Eye diameters and interdistances: AME 0.05; ALE 0.09; PME 0.06; PLE 0.08; AME–AME 0.07; AME–ALE 0.11; PME–PME 0.10; PME–PLE 0.19; AME–PME 0.07; ALE–PLE 0.05. Chelicerae paturon: length 0.75. Leg measurements: I: femur 1.20/ patella 0.48/ tibia 1.04/ metatarsus 0.88/ tarsus 0.56/ total 4.16; II: 1.10/ 0.54/ 0.75/ 0.92/ 0.54/ 3.85; III: 1.00/ 0.52/ 0.72/ 0.74/ 0.44/ 3.42; IV: 1.48/ 1.10/ 0.48/ 0.90/ 0.42/ 4.38. Leg formula 1243. Leg spination: metatarsus II–IV v1map. Copulatory ducts almost completely covered by the median field ( Fig. 12E ). Posterior edge of the epigynum round ( Fig. 12E ). Apex of the copulatory ducts bent ( Fig. 12F ). Horizontal spermathecae slightly curved ( Fig. 12F ).

Variation. Length (2♀): total 3.24–3.84; carapace 1.56–1.80; femur I 1.03–1.20.

Distribution. Only known from the type locality ( Fig. 27 ).

Figures

FIGURE 12. A–F. Goeldia goytacazes sp. nov. A–D: male, left palp; E–F: female epigynum: A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view.

FIGURE 16. A–F. Goeldia mirim sp. nov. A–D: male, left palp; E–F: female, epigynum: A. Ventral view; B. Tegular process, frontal view. C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view.

FIGURE 27. Distribution of Goeldia spp. in South America: Goeldia luteipes (square—purple); G. zyngierae (star—red) and G. gauderio (triangle—yellow); G. goytacazes (pentagon—orange); G. patellaris (circle—green). Type species are marked as black versions of each symbol when there are records to multiple localities.