Goeldia mexicana (O. Pickard-Cambridge, 1896 )

Almeida-Silva, Lina M. & Brescovit, Antonio D., 2024, Unraveling the mysteries of Goeldia Keyserling, 1891: revision, description of seven new species and first record from USA (Araneae: Titanoecidae), Zootaxa 5428 (2), pp. 151-193 : 174-175

Goeldia mexicana (O. Pickard-Cambridge, 1896)

( Figs 15 ; 24 )

Temecula mexicana O. Pickard-Cambridge, 1896: 170 , pl. 22, fig. 8 (holotype male from Teapa (17°32’07”N 92°45’03”W),

Tabasco, Mexico, deposited at NHM, examined); F.O. Pickard-Cambridge 1902: 359, pl. 83, figs 23–24 (descr. female). Goeldia funesta : Lehtinen 1967: 236 (syn). Goeldia mexicana : Leech 1972: 104, figs 185–186, 383–384 (removed from syn. with G. funesta ); World Spider Catalog

2023.

Remarks. Leech (1972: 104) mistakenly mentions that Lehtinen (1967: 236) synonymized G. luteipes and G. tizamina with Temecula mexicana .

Additional Material Examined. MEXICO. Veracruz: Emiliano Zapata, El Aguaje (19°25’00”N 96°37’00”W), 7 Km S of La Bandera, 1♀, 23.XII.1976, J. Reddel coll. ( AMNH) ; Catemaco, Lago Catemaco (18°24’N 95°07’W), 1♀, W. J. Gertsch & W. Ivie coll. ( AMNH) . Puebla: Grutas Atepolihuit , 5 km SE of Ceutzalan (20°01’07”N 97°31’16”W), 1♀, 18.XII.1976, D. McZengie & C. Soileau coll. ( AMNH) . Chiapas: Toniná (16°58’49”N 92°17’17”W), 15 Km E of Ocosingo, elev. 920 m, 1♀, 28.VI.1950, C. and M. Goodnight & L.J. Stannard coll. ( AMNH) ; San Cristobal de Las Casas (16°44’12”N 92°38’18”W), 1♀, C. and M. Goodnight coll. ( AMNH) ; El Real (16°54’N 91°45’50”W), Postrero, 3♀, 30.VI.1950, C. and M. Goodnight coll. ( AMNH) ; 6.4 Km SE of San Cristobal (16°42’N 92°36’W), 1♀, 23.VII.1966, J & W. Ivie coll. ( AMNH) ; 24 km NO of Arriaga , (16°25’N 94°01’W), pine forest, 1♀, 27.VII.1966 J. & W. Ivie coll. ( AMNH) . Tabasco: near Grutas de Cocona (17°34’7”N 92°56’37”W), 1♀, 25.VIII.1972, J. Cook, R. Mitchell & F. Russell coll. ( AMNH) ; Villahermosa (17°59’13”N 92°55’10”W), 1♀, 2.IV.1969 ( AMNH) . Oaxaca: San Felipe (18°04’00”N 96°31’59”W), 2♀, 24.VIII.1947, H. Wagner coll. ( AMNH) ; Orizaba (18°51’0”N 97°6’0”W), elev. 1200 m, 1♀, 7.VII.1941, H. Dybas coll. ( AMNH) . HONDURAS. Copán: Copán Ruins (14°50’N 89°09’W), 1♀, 7.III.1939, R. V. Chamberlin coll. ( AMNH) . COSTA RICA. Puntarenas: Monteverde (10°18’51.32”N 84°49’30.08”W), 12.8 Km of Belo M. V., (10°21’N 84°48’W), 1♀, Roth-Schroepfer coll. ( AMNH) . PANAMA. Chiquirí: South of Vulcano Baru (8°48’29”N 82°32’35”W), elev. 1000–1700 m, 1♀, 28.I.1987, E.S. Ross coll. ( CAS 9025932 ) ; Panama: Panama city (8°59’0”N 79°31’0”W), 1♀, II.1945, C.D. Michener coll. ( AMNH) .

Diagnosis. Goeldia mexicana differs from the other species by the male palp with tegular process triangular, short, and distally narrowed ( Fig. 15A ); and patellar apophysis straight, triangular, with tip rounded ( Fig. 15B–C ). The female epigynum can be recognized by its median field well developed, resembling G. luteipes but differing by the median area posteriorly projected ( Fig. 15E ); apex of the copulatory ducts dorsally projected, with enlarged bases and vertical spermathecae short ( Fig. 15F ).

Description. Male (AMNH). Coloration: carapace brown; chelicerae reddish brown; labium, endites, legs and sternum yellowish brown; opisthosoma pale brown. Total length: 5.56. Carapace: length 2.45, width 1.75, height 0.63. Opisthosoma: length 2.69, width 1.44. Clypeus: height 0.2. Eye diameters and interdistances: AME 0.06; ALE 0.09; PME 0.07; PLE 0.09; AME–AME 0.11; AME–ALE 0.13; PME–PME 0.16; PME–PLE 0.2; AME–PME 0.11; ALE–PLE 0.06. Paturon: 1.13 long. Leg measurements: I: femur 2.04/ patella 0.72/ tibia 1.80/ metatarsus 1.6/ tarsus 0.88/ total 7.04; II: 1.75/ 0.78/ 1.56/ 1.53/ 0.72/ 6.34; III: 1.64/ 0.8/ 1.28/ 1.4/ 0.64/ 5.76; IV: 1.93/ 0.75/ 1.75/ 1.56/ 0.63/ 6.61. Leg formula: 1423. Leg spination: femur I p1ap; metatarsus I v1r-1r- 1m, r1ap; II v1p-2- 1m, r1ap, p1ap; III v0-2-2ap, p1ap, r1ap; IV v2-2-1m-2ap, r0-2-0-1ap, p0-1-0-2ap. Male palp with apex of MLT partially sclerotized ( Fig. 15C ). Cymbium with modified setae ( Fig. 15D ). Pars pendula thickening subdistal ( Fig. 15D ).

Female (AMNH). Coloration: carapace, endites and chelicerae brown; labium dark brown; legs and sternum pale brown. Total length: 4.50. Carapace: length 2.0, width 1.6, height 0.72. Opisthosoma: length 2.5, width 1.9. Clypeus: height 0.14. Eye diameters and interdistances: AME 0.10; ALE 0.10; PME 0.08; PLE 0.09; AME–AME 0.07; AME–ALE 0.01; PME–PME 0.13; PME–PLE 0.19; AME–PME 0.10; ALE–PLE 0.08. Chelicerae paturon: length 0.89. Leg measurements: I: femur 1.54/ patella 0.66/ tibia 1.28/ metatarsus 1.22/ tarsus 0.64/ total 5.34; II: 1.34/ 0.69/ 1.12/ 0.98/ 0.58/ 4.72; III: 1.25/ 0.65/ 0.98/ 0.85/ 0.50/ 4.23; IV: 1.45/ 0.73/ 1.23/ 1.00/ 0.50/ 4.90. Leg formula: 1423. Leg spination: femur I p1sa; metatarsus I v0-1r-1, r1ap; II p1ap, r1ap, v1ap or v0-1r- 1m; III p1ap, r1ap, v1ap; IV v1m-0-1map or v0-1p-1map, p1ap, r1ap. Epigynum median field wide and covering part of the copulatory openings ( Fig. 15E ). Epigynal rim very narrow ( Fig. 15E ). Fertilization ducts close together, with 1/3 the width of the middle field ( Fig. 15E ). Copulatory ducts with enlarged base and heavily sclerotized ( Fig. 15F ). Vertical spermathecae very short, almost the height of the copulatory duct base ( Fig. 15F ). Apex of the copulatory duct dorsally bent ( Fig. 15F ). Apex of the horizontal spermathecae ventrally bent ( Fig. 15F ).

Variation. Coloration can be darker, with variable pattern where leg segments are proximally and/or dorsally lighter colored than the other segments/surfaces. Length (10 ♀): total 3.5–5.93; carapace 1.52–2.43; femur I 1.05– 1.81.

Distribution. Southern Mexico, Honduras, Costa Rica and Panama ( Fig. 24 ).

Figures

FIGURE 15. A–F. Goeldia mexicana (O. Pickard- Cambridge, 1896). A–D: male left palp; E–F: female epigynum: A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view.

FIGURE 24. Distribution of Goeldia spp. in North, Central and northern South America: Goedia funesta (star—purple); G. mexicana (triangle—red); G.chinipensis (pentagon—yellow). Type species are marked as black versions of each symbol, when there are records to multiple localities.