Mitella makinoi var. makinoi

M. Wakabayashi, 2006, Tiarella and Mitella, Flora of Japan, Volume 2 b: Angiospermae-Dicotyledoneae: Archichlamydeae (b), Kodansha, pp. 70-75 : 73

var. makinoi (H.Hara) Wakab.

in Acta Phytotax. Geobot. 28: 118 (1977).

Mitella makinoi H.Hara in J. Jap. Bot. 14: 519 (1938).

Mitella japonica auct. non Maxim., nec Miq.: Makino,

Icon FI. Jap. 1(2): 7 (1902).

Japanese name: Shikoku-charumeru-so.

Radical leaves ovate, cordate, acute or subacute, 2-7 cm long, 2-6 cm wide, 5-7 lobulate with acute tips or obsoletely lobulate, irregularly toothed. Flowering stems 20-40 cm tall. Flowers April to May. Calyx tube obconical; calyx lobes triangular-ovate, acute, often sparsely glandular along margin, erect. Petals (3-)5-pinnatiparted, spreading or reflexed, ca. 3.5 mm long; pinnae obliquely ascending, glandular dotted. Stamens 0.9-1.2 mm long. Styles 2, erect, ca. 0.8 mm long; stigmas thickened, shallowly or obscurely 2-4-lobed.

Chromosome number: 2n = 28 (Wakabayashi 1973).

Japan: Shikoku and S. Kyushu (Kagoshima and Kumamoto Pref.). Moist, shady places along streams in mountainous areas. Endemic.

Icones: Makino, Icon. FI. Jap. 1(2): t. 4 & 5 (1902), as M. japonica ; Nakai & Honda, Nov. FI. Jap. 3: f. 36 (1939); Kitamura & Murata, Herb. PI. 2: f. 72 4, as M. makinoi ; Satake et al., Herb. PI. 2: photo. 152 3 & 4; Makino, rev. ed.: f. 972, saltern pro parte, as M. japonica .