Ipomoea dubia Roem. & Schult.

Wood, John R. I., Munoz-Rodriguez, Pablo, Williams, Bethany R. M. & Scotland, Robert W., 2020, A foundation monograph of Ipomoea (Convolvulaceae) in the New World, PhytoKeys 143, pp. 1-823 : 1

326. Ipomoea dubia Roem. & Schult. , Syst. Veg. 4: 216. 1819. (Roemer and Schultes 1819: 216)

Ipomoea angulata Ortega , Nov. Rar. Pl. Dec. 7 -8: 83 1797. (Ortega 1797 -1800: 83), nom. illeg., non Ipomoea angulata Lam. (1793) . Type. Plants grown at Madrid from seed sent by Ruiz and Pavón (lectotype OXF00006441, designated here; isolectotype P).

Quamoclit ruiziana G. Don , Gen. Hist. 4: 258. 1838. (Don 1838: 258), nom. illeg., superfl., Type. Based on Ipomoea angulata Ortega

Type.

Based on Ipomoea angulata Ortega

Description.

Prostrate, ascending or erect annual herb; stems glabrous or pubescent. Leaves petiolate, 2-7 × 1.5-5.5 cm, entire (rarely shallowly 3-lobed), ovate, cordate with rounded or angled auricles, apex finely acuminate, mucronate, margin entire or (rarely) undulate, adaxially glabrous, abaxially glabrous or pubescent, especially on the veins; petioles 2-7 cm, glabrous or pubescent. Inflorescence of long-pedunculate, few-flowered axillary cymes; peduncles (1.4-)3-7.5 cm, remaining erect in fruit, glabrous or pubescent; bracteoles 1-3 mm, lanceolate, acuminate; secondary peduncles 5-7 mm; pedicels 2-7 mm, often angled, glabrous or pubescent; sepals unequal, outer ovate with scarious margins, midvein sometimes extended to form a wing, glabrous or puberulent, 4-5 × 2-3 mm, terminating in a mucro 4-8 mm long, inner sepals 5-6 × 3 mm with a mucro 4-8 mm long; corolla 2-2.5 cm long, hypocrateriform, scarlet, glabrous, limb c. 1.5 cm diam., unlobed. Capsules 5-6 × 6-7 mm, compressed globose, rostrate, the persistent style c. 3 mm long, pubescent or glabrous; seeds 4 × 2 mm, distinctly tomentose.

Distribution.

Endemic to Peru and Ecuador between 400 and 2700 m, most records from coastal and lower western semi-desert slopes of the Andes in the Lima area.

PERU. Ancash: E. Cerrate et al. 5180 (MO, USM). Cajamarca: Contumaza, A. Sagástegui & López 9166 (FTG, MO), 10529 (FTG, MO), 1573 (F). Ica: Mun. Yauca del Rosario, O. Whaley et al. 460 (K). La Libertad: P. Nuñez et al. 6265 (CUZ); Trujillo, A. Sagástegui & Cabanillas 8743 (TYG, HUT); ibid., A. Sagástegui & J. Mostacero 10447 (MO); ibid., Contumaza, El Balconcito, A. Sagástegui & S. Leiva 16404 (OXF). Lambayeque: E. Cerrate et al. 5242 (USM). Lima: S.G.E. Saunders 855 (K); C.A. Weatherby 11320 (K); A. Gentry et al. 19912 (FTG, MO); entre Chosica y Surco, R. Ferreyra 6938 (MO, USM). Piurá: Chililique, Bajo Naranjo, E. Laure 5477 (P); Huancabamba, La Beatita, Llatas Quiroz 2455 (F). San Martín: Chrostowski 69-197 (S). Tumbes: Zarumilla, Lechugal, R. Ferreyra 10659 (MO, USM).

ECUADOR. Chimbarazo: Cañon del Río Chanchan, Huigra, W.H. Camp 2970 (FTG, S); Huigra, J.N. & G. Rose 22298 (NY, US). Loja: Sabanilla, C. Quintana et al. 2887 (QCA). Pichincha: Reserva Pululahua, Canton Quito, C. E. Cerón 2258 (MO). Tungurahua: J.E. Madsen 36442 (AAH).

Typification.

There appears to be no syntype at Madrid so we have selected the specimen at OXF as the lectotype as this is a more complete specimen than that at Paris.

Note.

Distinct because of the relatively large sepals with long erect awn-like mucros. The capsule is very unusual, being often pubescent. The short, angled or winged pedicels are also noteworthy.