Phlaeopterus loganensis Hatch, 1957

Mullen, Logan J., Campbell, J. M. & Sikes, Derek S., 2018, Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini), The Coleopterists Bulletin 72, pp. 1-1 : 1-

13. Phlaeopterus loganensis Hatch, 1957 (Figs. 4A, 12A, 16H, 24C, 29E)

Phlaeopterus loganensis Hatch 1957: 60 [original description]. Campbell and Davies 1991: 5; Bousquet et al. 2013: 89.

Type Locality. Logan Pass , Glacier National Park, Montana, USA .

Redescription. Habitus: Length 5.8–7.4 mm. Dark brown to black; head often black, pronotum and elytra lighter; antennae, palpi, and legs even lighter, legs becoming progressively lighter from base to apex (Fig. 4A). Head: Width across eyes to head length subequal to slightly wider than long. Interantennal groove broadly and shallowly impressed. Anteocellar foveae large, deeply impressed. Eyes glabrous or with less than 10 scattered setae near ventral margin. Antennomeres 5–10 at least 2 times longer than wide; antennomeres 4–11 each with many sensory pits with papilliform projections. Ocelli present. Nuchal constriction vague. Mandibular molar area with L-shaped row of setae. Labrum with sensory pores along entire surface. Labial palpi with 3 rd palpomere 1.6–1.8 times longer than palpomere 2. Thorax: Pronotum moderately wide ( Fig. 16H ), length to width ratio = 0.68–0.75; ratio of pronotal width to head width = 1.52–1.67; maximum width subequal to elytral width at humeral angle; punctures separated by distance slightly less than diameter of a puncture on center of disc, almost contiguous towards margin; lateral margins explanate posterad but not anterad lateral foveae; lateral foveae deeply impressed. Elytra with humeral angles convex; epipleural carina not projecting; 2.2–2.7 times longer than pronotum; apical margins broadly triangular, elongated at suture ( Fig. 29E ). Wings fully developed in most individuals, brachypterous in very few. Mesosternum with projecting tooth; longitudinal carina along midline of mesosternum indistinct. Legs: Apices of all tibiae without pubescence, length of subglabrous apex of metatibia as ratio to metatibial length in males = 10.2–17.3, in females = 8.1–13.0. Metatrochanter without tooth on apical margin. Abdomen: Wing-folding spicules on tergites IV and V broad, narrowly separated; wing-folding patches on tergite VI small, circular to slightly transverse. Apical palisade fringe on tergite VII present. Aedeagus: Length 1.40–1.55 mm long. Median lobe narrow, elongate, lateral margins constricted and subparallel near midline, expanding anterad and posterad midline ( Fig. 24C ). Parameres narrow, convexly curved. Internal sac elongate, lightly sclerotized, dorsal half evenly, lightly covered in microspinules.

Type Specimens. Holotype male (UAMObs: Ento:235773) and allotype female (UAMObs: Ento:235774) labeled as follows: G.N.P. Mont., Logan Pass, Aug. 22, 1939, M.H. Hatch/ TYPE ³

Bionomics. Adults have been collected during June–September at 600–2,400 m elevation in wet moss and piles of rocks at the edges of fast, cold streams, in the splash-zone of waterfalls and cascades, and under rocks at the edges of snowfields or snowmelt ponds. Specimens collected at low elevations were associated with springs or fast streams that remain cold to lower elevations.

Remarks. Phlaeopterus loganensis can be distinguished from all other Phlaeopterus species by having the apices of the elytra triangular in both the male and the female ( Fig. 29E ). Although P. elongatus and P. filicornis have the elytral apices triangular in the female, no other species of the genus has the elytral apices triangular in both sexes. Phlaeopterus loganensis is most similar to P. olympicus , and these two species can be distinguished by characters discussed in the Remarks section of the latter species.

Figures

Fig. 16. Pronota, dorsal view, of Phlaeopterus species. A) P. kavanaughi, B) P. castaneus, C) P. cavicollis, D) P.

Fig. 29. Phlaeopterus species. Right mandibles, ventral view, of A) P. obsoletus, B) P. bakerensis, C) P. elongatus. D) P. bakerensis, humeral angle of elytron, dorsal view. Apices of elytra of E) P. loganensis, F) P. hatchi, G) P. filicornis female.

Fig. 24. Phlaeopterus species, aedeagi (left = ventral view; right = dorsal view of median lobe). A) P. occidentalis, B) P. olympicus, C) P. loganensis, D) P. fusconiger.