Phlaeopterus kavanaughi Mullen and Campbell, 2018

Mullen, Logan J., Campbell, J. M. & Sikes, Derek S., 2018, Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini), The Coleopterists Bulletin 72, pp. 1-1 : 1-

11. Phlaeopterus kavanaughi Mullen and Campbell , new species

( Figs. 3C , 11A, 16A , 21E , 35A–B , 41B ) Zoobank.org/ urn:lsid:zoobank.org:act:6CE135BC-1612-4382-92FD-6F6FB45A6DBD

Type Locality. Lassen National Park , California, USA .

Description. Habitus: Length 6.2–7.6 mm. Dark brown to reddish brown to yellowish brown; head and abdomen nearly black, elytra usually distinctly lighter reddish to yellowish brown; pronotal margins often also distinctly lighter reddish brown to yellowish brown; palpi, antennae, and legs reddish brown to yellowish brown ( Fig. 3C ). Head: Moderately broad ( Fig. 35A–B ), ratio of width across eyes to head length = 1.0–1.3. Interantennal groove broadly and deeply impressed. Anteocellar foveae large, deeply impressed. Eyes glabrous or with less than 10 scattered setae near ventral margin. Antennomeres 5–10 at least 2 times longer than wide; antennomeres 4–11 each with many sensory pits with papilliform projections. Ocelli present. Nuchal constriction vague. Mandibular molar area with L-shaped row of setae. Hypopharynx as in Fig. 41B . Labrum with sensory pores along entire surface. Labial palpi with 3 rd palpomere 1.6–1.8 times longer than 2 nd palpomere. Thorax: Pronotum broad ( Fig. 16A ), length to width ratio = 0.71–0.79; ratio of pronotal width to head width = 1.49–1.67; maximum width subequal to elytral width across humeral angles; punctures on dorsal surface separated by average distance slightly greater than diameter of a puncture; lateral margins explanate posterad lateral foveae, not explanate anterad lateral foveae; lateral foveae deeply impressed. Elytra with humeral angles convex; epipleural carina not projecting; 2.1–2.6 times longer than pronotum. Wings fully developed in most individuals, brachypterous in very few. Mesosternum with projecting tooth; longitudinal carina along midline of mesosternum complete but not strongly carinate. Legs: All tibiae with dense pubescence from base to apex. Metatrochanter without tooth on apical margin. Abdomen: Wing-folding spicules on tergites IV and V broad, transverse, narrowly separated; tergite VI lacking wing-folding patches. Aedeagus: Length 1.16–1.34 mm. Median lobe approximately evenly, somewhat convexly narrowed from base to apex; with vague ventral longitudinal carina at apex ( Fig. 21E ). Parameres narrow. Internal sac elongate, tube-like, moderately sclerotized; with distinct median transverse fold.

Type Specimens. Holotype male (UAMObs: Ento:235218) and allotype female (UAMObs: Ento:235831) labeled as follows: CAL. Lassen N. P., base of Summit Tr., 8500’, 16.VII.1979, JM & BA Campbell/ HOLOTYPE ³ (or ALLOTYPE ♀) Phlaeopterus kavanaughi , desig. L.J. Mullen and J.M. Campbell, CNC No. 18464 (red label). Both specimens are deposited in the CNC. Paratypes (n = 183) are deposited in AMNH, NHMUK, CAS, CSCA, FMNH, MCZ, OSAC, USNM, and UCRC.

Distribution. Phlaeopterus kavanaughi is known from the Sierra Nevada and the Cascade Range of California north to Oregon ( Fig. 11A).

Bionomics. Adults have been collected during July–August at 1,830–3,400 m elevation, with a single specimen (UAMObs:Ento:234158) collected at 90 m in Multnomah County, Oregon. This species has been collected at night on snowfields, where

Figures

Fig. 3. Phlaeopterus species, habitus. A) P. hatchi, male (UAMObs:Ento:232429), B) P. houkae, male (UAMObs:

Fig. 16. Pronota, dorsal view, of Phlaeopterus species. A) P. kavanaughi, B) P. castaneus, C) P. cavicollis, D) P.

Fig. 21. Phlaeopterus species, aedeagi (left = ventral view; right = dorsal view of median lobe). A) P.

Fig. 35. Heads, dorsal and dorsal oblique views, of A–B) Phlaeopterus kavanaughi, C–D Phlaeopterus hatchi. Eyes, dorsal view, of E) Lesteva longoelytrata, F) Unamis sp. undescribed.

Fig. 41. Phlaeopterus species, hypopharynges, dorsal view, of A) P. obsoletus, B) P. kavanaughi, C) P. elongatus. D) Phlaeopterus lagrandeuri, metatrochanter with apical tooth. Heads, ventral view, of E) Lesteva pallipes, F) Lesteva pubescens.