Psammoecus dentatus Grouvelle, 1883

Yoshida 1, Takahiro, Karner, Michael & Hirowatari, Toshiya, 2018, A Revision of Taiwanese Species in the Genus Psammoecus Latreille (Coleoptera, Silvanidae), Zoological Studies 57 (18), pp. 1-18 : 12

Psammoecus dentatus Grouvelle, 1883

( Fig. 1D )

Psammoecus dentatus Grouvelle, 1883: 290 . Type locality: New Guinea. Type examined.

Psammoecus quadrimaculatus: Hirano 2009: 64 , 65, 82, fig. 4. - Hirano 2010: 12, 14. (misidentification)

Psammoecus scitus Yoshida and Hirowatari, 2014: 18 , figs. 1A, 3, 13A-C. Type locality: Japan; Type examined. - Yoshida and Hirowatari 2015: 92-94 . syn. nov.

Type series: Lectotype: male, “N. Guinea | Hatam Vll | Beccari 1875”; “SYNTYPUS | Psammoecus | dentatus | Grouvelle, 1882” ( MSNG; studied by MK in 2015 and TY in 2017, here designated) . Paralectotypes: 1 male and 1 female, same label data as lectotype. ( MSNG; studied by MK in 2015, here designated).

Specimens examined: JAPAN: 1 male, Kûra, Ishigaki Island, Okinawa Prefecture, 22-IX-2012, T. Yoshida leg. ( EUMJ; formerly holotype of P. scitus ). TAIWAN: [Hualien County] 1 ex., Liyuchih, 14-VIII-1966, H. Kamiya leg. ( KUM); 2 exs, same locality, 15-VIII-1966, H. Kamiya leg. ( KUM); 2 exs, Juisui (Naka spa) (= Ruisui Township), 27-XI-1962, K. Baba leg. ( EUMJ); 2 exs, Fu Yuan For. Rec. Area, 28-V-1997, C. W. & L. B. O’Brien leg. ( FSCA). [Nantou County] 1 male (Catalog No. NMNS ENT 1582-60), Fenghuangku, Lugu Township, 20-22- IX-1993, C. S. Lin & M. L. Chan leg., UV Light ( NMNS). [Chiayi County] 1 ex. (Catalog No. NMNS ENT 425-406), Yunghsing, Dapu Township, 15-XII-1988, K. W. Huang leg., Sweeping Net ( NMNS). [Taitung County] 2 males & 3 exs, Doulan, Donghe Township, 9-IV-2012, J. Yamasako leg. ( ELKU); 1 ex., Linchan, Daren Township, 6-VI-2013, S. Tsuyuki leg. ( ELKU); 5 exs, Jhiben Hot Spring, Beinan Township, 24-V-1969, S. Hisamatsu leg. ( EUMJ); 1 ex., Lutao (= Green Island), Green Island Township, 26-28-III-1998, M. Sato leg ( EUMJ); 3 exs, Island Lanyu (= Orchid Island), Lanyu Township, 8-IV-1978, Kôzô Murakami leg. ( EUMJ); 15 exs, Yung Hsin Farm, Lanyu Island (= Orchid Island), Lanyu Township, 29-30-VII-2008, Mendel & Barclay leg. ( BMNH; studied by MK). [Kaohsiung City] 1 ex., Kosempo (= Jiaxian District), I-1908, H. Sauter leg. ( ZMHB; studied by MK); 1 ex., Paiyunshan (= Baiyunshan), near Chiashien (= Jiaxian), Jiaxian District, 21-III-1977, Yutaka Notsu leg. ( EUMJ); 2 exs, Fa Kuo Shan, near Liu Kui (= Liouguei), alt. 800 m, 6-X-1986, K. Baba leg. ( EUMJ); 1 ex., Liu Kui (= Liouguei), 30- III-1986, K. Baba leg. ( EUMJ); 1 ex., same locality, 2-IV-1986, K. Baba leg. ( EUMJ); 1 ex., same locality, 4-VIII-1978, T. Niisato leg. ( EUMJ). [Tainan City] 1 ex., Kantoushan, Dongshan District, 17-II-2013, W. -C. Liao leg. ( TARI). [Pingtung County] 2 exs, Hengchun, 4-VIII-1966, H. Kamiya leg. ( KUM); 1 ex. (Catalog No. NMNS ENT 6215-77), Chungchienlu, Mudan Township, 27-28-VII-2009, W. T. Yang & K. W. Huang leg., UV Light Trap ( NMNS); 1 male & 1 female, Dong Yuan Forest Recreation Area, Dong Yuan Village, Mudan Township, Pingtung City, 10-VII-2014, T. Yoshida leg. ( ELKU); 2 exs (Catalog No. NMNS ENT 6259- 276 & 580), Chiupeng, Manzhou Township, 19-IV- 1-V-2009, M. L. Jeng & T. J. Chen leg., Flighintercept Net. ( NMNS); 2 exs, Kenting, 4-IX-1983, J. B. Happner leg. ( FSCA); 1 ex., Kenting Park, 17- VIII-1968, M. Nishikawa leg. ( EUMJ); 1 ex., same locality, 17-III-1977, Yutaka Notsu leg. ( EUMJ).

Diagnosis: This species can be distinguished from other Taiwanese Psammoecus species by the small, even-sized lateral teeth on its pronotum and the irregular distances between the adjacent teeth.

Distribution: Taiwan; Japan; New Guinea.

Remarks: This species was described as P. scitus by Yoshida and Hirowatari (2014).

This study examines many specimens in this species, including P. dentatus and P. scitus type specimens. We found no significant differences between these two species in their morphology, including their male genitalia; thus, P. scitus is synonymized with P. dentatus .

Specimens from an extremely large area have been studied in the course of related revisionary work. Psammoecus dentatus may be recorded from many more South East Asian localities.

Figures

Fig. 1. Psammoecus species of Taiwan, habitus. (A) P. taiwanensis sp. nov. (B) P. delicatus Grouvelle (C) P. harmandi Grouvelle (D) P. dentatus Grouvelle (E) P. trimaculatus Motschulsky (F) P. triguttatus Reitter (G) P. labyrinthicus Yoshida and Hirowatari (H) P. simonis Grouvelle (I) P. hiranoi Yoshida and Hirowatari. Scale bar = 1.0 mm.