Toktokkus vialis, (HAAG-RUTENBERG)

Kamiński, Marcin J., Gearner, Olivia M., Kanda, Kojun, Swichtenberg, Kali, Purchart, Luboš & Smith, Aaron D., 2021, First insights into the phylogeny of tok-tokkie beetles (Tenebrionidae: Molurina, Phanerotomeina) and examination of the status of the Psammodes vialis species-group, Zoological Journal of the Linnean Society 191, pp. 883-901 : 899

TOKTOKKUS VIALIS SELLATUS (HAAG- RUTENBERG)

= Psammodes kuisup Péringuey, 1908 , synon. nov. (as P. ‘ kuisip ’ in Gebien, 1937 and Kaminski et al., 2019a). = Psammodes sellatus uriai Koch , synon. nov.

Justification: When describing Psammmodes kuisup (in erratum, referred to in the description as Psammodes tuberculipennis ), Péringuey (1908) was apparently unaware of Psammodes sellatus Haag-Rutenberg and, therefore, did not provide diagnosable characters to differentiate the two species. Examination of specimens, including the P.sellatus holotype and a specimen collected at the type locality and determined by Koch as P. kuisup , and comparison of the species descriptions, reveal no easily diagnosable characters between these two species.

Koch (1953b) described the subspecies P. sellatus uriai as differing from P. sellatus sellatus by the following characters: a more elongate body, a slenderer pronotum with less prominent punctation and tubercles less dense and organized in rows (as opposed to denser and more irregular in sellatus ). However, when comparing specimens, these differences appear minor and not sufficient to justify classifying these as separate subspecies.

NOTES ON PHANEROTOMEINA :

GENUS CHILIARCHUM KOCH ( SEPIDIINI : PHANEROTOMEINA )

l s i d u r n:l s i d:z o o b a n k.o r g:a c t: C B7 1 2 3 7 1-B6 0 4- 4D1B-93A2-1258C6920572

Type species: Moluris (Phanerotoma) bertolonii Guérin-Méneville (by original designation).

Diagnosis: From Koch (1954): submarginal area of the sides of the pronotum, and often that of the elytra, with either a broad stripe of white to yellow subtomentose hairs along the lateral carina or with densely conglomerated micropunctation or granulation with associated scattered short, fine, squamulated bristles; pronotum with two prebasilar impressions at or near the posterior angles, filled with pale subtomentose hairs or densely micropunctured with scattered short, fine bristles; males with a reddish brown brush of hairs on the outer edge of the upper surface of the mesotibiae, sometimes extending to the tibial base.

Species and subspecies included (8): Chiliarchum arnoldi arnoldi ( Koch, 1952) comb. nov., C. arnoldi sabianus ( Koch, 1952) comb. nov., C. bertolonii (Guérin-Méneville, 1844) comb. nov., C. freyi ( Koch, 1952) comb. nov., C. guerini guerini (Haag- Rutenberg, 1871) comb. nov., C. guerini lawrencii ( Koch, 1954) comb. nov., C. guerini mancus ( Koch, 1954) comb. nov., C. junodi (Péringuey, 1899) comb. nov.