Harmonides Kirkaldy

McKamey, Stuart H., Wallner, Adam M. & Porter, Mitchell J., 2015, Immatures of the New World treehopper tribe Amastrini (Hemiptera, Membracidae, Smiliinae) with a key to genera, ZooKeys 524, pp. 65-87 : 73-74

Taxon classification Animalia Hemiptera Membracidae

Harmonides Kirkaldy Figs 24-25, 52-53

Diagnosis.

Head and pronotum without scoli; meso- and metanota and terga III-VIII each with short paired scoli; body evenly covered with chalazae with long setae.

Nymphal description.

Overall body. Chalazal setae long; dorsal contour of abdomen in lateral view curvilinear; scoli splayed away from each other. Head. Scoli present or absent; scoli, if present, directed dorsad; chalazal setae simple, hairlike. Prothorax. Premetopidium scoli present or absent; premetopidium scoli, if present, directed anteriorly, and about 2-4 5 basal width; postmetopidium scoli absent; posterior extension of pronotum surpasses anterior margin of metanotum. Mesothorax. With paired scoli dorsally; scoli bearing tuberculate chalazae or without chalazae; scoli directed anteriorly or dorsally; forewing pad costal margin straight; forewing pad chalazae sparsly, their setae short; dorsal scoli 2-4 5 basal width; forewing pad without costal chalazae. Metathorax. With paired scoli dorsally, directed dorsally or almost so, 2-4 5 basal width, bearing tuberculate chalazae. Legs. Chalazae of tibia on both lateral margins but with or without chalazae on dorsal surface also. Abdomen. Terga III-VIII ventrolateral margins with single enlarged chalaza or without enlarged chalazae; terga III-VIII dorsal scoli all subequal in size or size decreasing posteriorly, tallest dorsal scoli 2-4 5 basal width; tergum IV dorsal scoli directed dorsally or almost so; terga III-VIII lateral rows not manifested; tergum III with paired apically acute scoli dorsally. Segment IX. Dorsal length subequal to combined length of segments V-VIII or VI-VIII; without dorsal enlarged chalazae or scoli at apex; ventral extension subequal to dorsal extension.

Material examined.

Harmonides reticulata (Fabricius), 2 nymphs, 1 adult, VENEZUELA: Estado Zulia, Dist. Pirija, Tocuco, 28-29 June 1984, S.H. McKamey lot #680. (USNM). Harmonides sp. 1, 1 adult, 1 nymph, PANAMA, Barro Colorado Island, Canal Zone,VII-VIII 1942, JasZetel No. 4485 (USNM).

Distribution.

Brazil northward to Mexico.

Discussion.

Harmonides reticulata is a common widespread species that SHM has often found in aggretations of adults and nymphs. Harmonides sp. 1 is curious in that it has delicate premetopidial scoli (Fig. 25) and therefore resembles Neotynelia . The two genera differ in the presence, in Neotynelia , of scoli on the head as well.