Helictophanes flava Muadsub and Pinkaew, 2014

Muadsub, Sopita & Pinkaew, Nantasak, 2014, Two new species of Enarmoniini (Lepidoptera: Tortricidae) from eastern Thailand, Zootaxa 3841 (1), pp. 127-134 : 128-130

Helictophanes flava Muadsub and Pinkaew , n. sp.

( Figs. 1A–B , 2A, 2C , 3A , 4A )

Diagnosis. This species is characterized by the following features: forewing yellow with a dark triangular dorsal mark and an irregular, curved band from the middle of the costa to the termen; hindwing veins M 3 and CuA 1 long stalked; male hindwing with a long ribbon-shaped anal lobe nearly reaching the anal angle; and short, nearly straight labial palpi with a small, very short terminal segment. The male genitalia are characterized by a narrow apex of the tegumen with dense scale sockets laterally; and by a valva with a distinct, curved neck and an elongate elliptical cucullus with a very short apical spine. The female genitalia are characterized by a ring-shaped sterigma, incomplete posteriorly; a very short ductus bursae; and a corpus bursae with two large triangular, bladelike signa with elongate hollow bases and a sclerotized plate posteriorly. H. flava , n.sp. differs from previously described species of Helictophanes by the presence of a sclerite on the posterior area of corpus bursae.

Description. Head. Ocellus large, vertex covered with golden yellow scales, upper frons golden yellow, with pale yellow, appressed scales on lower frons; labial palpus rather short (1.5X diameter of eye) ( Fig. 2A ), porrect, yellow, with brown to dark brown scales dorsobasally on second segment, first segment short, second segment long and nearly straight, gradually widening toward apex, terminal segment short, apex acute, pointing downward; antenna to beyond middle of forewing, golden yellow.

Thorax. Smooth, without raised scale tuft posteriorly, light brown; legs unmodified. Forewings subrectangular, length 2.9-3.0 mm in male (n = 2) ( Fig. 1A ) 3.8–3.9 mm in female (n = 2) ( Fig. 1B ); without costal fold in male, costa curved near base and before apex, apex rounded, termen oblique and nearly straight; forewing venation with all R veins separate, R 5 reaching to termen below apex, M 3 and CuA 1 parallel and very close in basal half ( Fig. 2C ); ground color pale yellow, with six dark brown spots along costa, three basally and the other three distally, and a long narrow dark brown band medially, curved and extending from costa to termen at M 1, brown to dark brown; dorsum with large, triangular brown mark medially, edged with dark brown scales; underside pale brown, speckled paler reflecting wing pattern. Hindwing narrow, apex pointed, with long ribbon-shaped anal lobe in male, nearly reaching anal angle; wing venation with M 3 and CuA 1 connate to long-stalked, without trace of CuP, 3A extending along basal half of anal margin, half as long as 1A+2A ( Fig. 2C ); upper side translucent, brown, paler towards base; underside pale brown.

Abdomen. Male genitalia ( Fig. 3A ). Tegumen moderately high, weakly sclerotized, dorsolaterally with narrow lobe with dense sockets; socius elongate membranous, sparsely covered with hairs; gnathos represented by a weak sclerotized band, arising from dorsal 1/3 of tegumen; vinculum small; valva moderately sclerotized, with large basal excavation, costa curved; sacculus large, subrectangular, with moderately dense long setae apicoventrally; with deep ventral excavation; cucullus long, slender, elliptic to band-shaped, slightly widening beyond neck, rounded apically with small, single dorsoapical spine; juxta small; caulis rather long; phallus cylindrical, straight, moderately long and wide, slightly tapering to apex; with numerous lanceolate cornuti, nearly as long as phallus. Female genitalia ( Fig. 4A ). Papillae anales with dense setae. Tergum VIII smooth except for lateral triangular extensions with sparse scale sockets; sternum VII weakly sclerotized, moderately scaled, sterigma beyond posterior margin of sternum VII, subcircular ridge, wider laterally, with dense microtrichia around ostium; colliculum small, irregular ring, moderately sclerotized; ductus bursae short and wide, ductus seminalis arising from lobe near bursae neck with elongate and sinuate sclerite on posterior area end of corpus bursae; corpus bursae subovate with two large, unequal signa: triangular blades with long hollow bases.

Holotype ♂. Thailand: Trat Prov.: Trat Agroforestry R.St., 12°23'43"N 102°40'32"E, ca. 30 m, 12 Oct 2012, N. Pinkaew, np5396 (genitalia slide NP2016). Deposited in BMNH.

Paratypes. Thailand: Trat Prov.: Trat Agroforestry R.St., 12°23'43"N 102°40'32"E, ca. 30 m, 12 Oct 2012, N. Pinkaew (1♂, genitalia slide NP2014, 2♀, genitalia slide NP2012, NP2017). Deposited in KKIC and THNM .

Etymology. The specific epithet flava (latin = yellow) refers to the ground color of forewing.

Distribution. Thailand (Trat).

Figures

FIGURE 1. Adults of Helictophanes and Cyphophanes. A. Helictophanes flava, n. sp. (holotype male). B. H. flava, n. sp. (paratype female). C. Cyphophanes khitchakutensis, n. sp. (holotype male). D. C. khitchakutensis, n. sp. (paratype female) (scale bar=2 mm).

FIGURE 2. Labial palpi and wing venation. A-B. Labial palpi; A. Helictophanes flava, n. sp. B. Cyphophanes khitchakutensis, n. sp. C-D. Wing venation; C. H. flava, n. sp. D. C. khitchakutensis, n. sp.

FIGURE 3. Male genitalia. A. Helictophanes flava, n. sp. (holotype male). B. Cyphophanes khitchakutensis, n. sp. (holotype male).

FIGURE 4. Female genitalia. A. Helictophanes flava, n. sp. (paratype female). B. Cyphophanes khitchakutensis, n. sp. (paratype female).