Anillini Jeannel, 1937

Harden, Curt W. & Caterino, Michael S., 2024, Systematics and biogeography of Appalachian Anillini, and a taxonomic review of the species of South Carolina (Coleoptera, Carabidae, Trechinae, Anillini), ZooKeys 1209, pp. 69-197 : 69-197

Tribe Anillini Jeannel, 1937

Adult diagnosis.

In the United States, adult specimens of Anillini are the only carabids that are both eyeless and possess subulate palpomeres.

Larval diagnosis.

Late-instar larvae of Anillini in South Carolina share the following characters: body pale and soft, largely unsclerotized (Fig. 13 A, B ); legs with single tarsal claw; stemmata absent; coronal suture absent; retinaculum large; penicillus present and consisting of several setae; urogomphus with seven large setae (Fig. 14 A, E ). Other known carabid larvae in SC of similar size with a single tarsal claw and potentially lacking stemmata are the genera Trechus , Semiardistomis , Clivina , and the tribe Tachyini . Larvae of Trechus are easily recognized by the possession of 4 - segmented labial palps and 5 - segmented maxillary palps (versus 2 - and 4 - segmented, respectively, in anillines). Semiardistomis and Clivina can be readily separated by the small size of the retinaculum and a penicillus consisting of a single, large seta. South Carolina anilline larvae are similar to larvae of the tribe Tachyini , but differ by having seven large setae on the urogomphus (Fig. 14 E ), versus six in tachyines.

Figures

Figure 13. Dorsal habitus of late-instar larvae A Serranillus dunavani (Jeannel) B Anillinus jancae. Scale bars: 1 mm.

Figure 14. Details of late-instar larvae of Appalachian Anillini A head of Anillinus jancae sp. nov., dorsal aspect, left antenna and right mandible omitted B left mandible of Serranillus dunavani (Jeannel), dorsal aspect C right maxilla of A. jancae, dorsal aspect D right maxilla of S. dunavani, dorsal aspect E urogomphi of A. jancae, dorsal aspect. Scale bar: 0.1 mm.