Cheumatopsyche chitawana Oláh & Johanson, 2008

Oláh, J. & Johanson, K. A., 2008, Generic review of Hydropsychinae, with description of Schmidopsyche, new genus, 3 new genus clusters, 8 new species groups, 4 new species clades, 12 new species clusters and 62 new species from the Oriental and Afrotropical regions (Trichoptera: Hydropsychidae), Zootaxa 1802, pp. 1-248 : 205

Cheumatopsyche chitawana Oláh & Johanson , new species

Fig. 341–344

Cheumatopsyche chitawana belongs to the Cheumatopsyche ningmapa , new species cluster. Here we separate this new species cluster as characterized by slender and highly sinuate harpagones, reduced mesoapical smooth lobe and shortened apicoventral setose lobes with knob-like apex. The species cluster contains totally 4 species: C. chitawana , C. ningmapa Schmid from Bhutan, C. naumanni Malicky from Afghanistan, Pakistan and Nepal, and C. charites Malicky & Chantaramongkol from Thailand, Laos, Vietnam and Malaysia ( Oláh et al. 2008). Cheumatopsyche chitawana is easily recognized by having knob-like apex of the apicoventral setose lobes not produced. Instead they originate posteriorly on segment X. As a result, the dorsal interlobular gap is shallower. Furthermore, segment X broadening before apex due to the longitudinal and lateral rim of the apicoventral lobes, easily seen in dorsal view.

Male. Body and wings pale brown with lighter pubescence; legs and antennae stramineous. Maxillary palp formula I-(III, IV)-II-V, segment V as long as sum of segments I–IV. Head dorsum pale brown, with 9 pale brown setal warts. Swollen setal wart absent on each proepisternum. Setal wart present on each precoxale. Pretarsal claws asymmetrical, laterally flanked by setal bundle. Mid- and hind-leg claws asymmetrical.

Wings. Forewing evenly pale, with slightly darker veins. Forewing crossveins m-cu and cu almost meeting in oblique line. Cu2 and A1 running separately before costa. Hind wing Sc and R meeting slightly posteriorly of crossvein r. Crossveins r and s tangential. Fork 1 absent. Forewing length 7.0 mm.

Male genitalia. Abdominal segment IX fused annularly, tergum as long as sternum ( Fig. 341). Anterior margins of segment IX regularly bow-shaped, rather flat dorsally in lateral view ( Fig. 341). Apical lobe on posterolateral margins blunt right-angled, located above less sclerotized articulation cavity of gonocoxites. Spine row on posterior margin of segment IX intermittent, heterogeneous. Spines on dorsolateral spiny lobes 2 times longer than spines on apical lobes; in dorsal view dorsolateral spiny lobes protruding and surrounding deep, wide, concave indentation or diverticulum ( Fig. 342). Inter-segmental step between segments IX and X pronounced, deep, sharply angled. Segment X long, nearly triangular in lateral view ( Fig. 341); regularly quadrangular in dorsal view ( Fig. 342); broadening apical half produced by ventral rim of apicoventral setose lobes; bilobed in dorsal view ( Fig. 342). Dorsal interlobular gap wide and shallow. Sutures well developed on both sides, transverse, crossing segment X obliquely; longitudinal sutures continuing from apicoventral setose lobes; meeting transverse suture into Y. Smooth dorsomesal plate reduced, ending in concavity. Mesal part of apicoventral setose lobes fusing with segment X; apex free, blunt, knob-like, curving slightly mesad and pressed into body of segment X ( Fig. 342); setae restricted to apical tip. Lateral setose areas (superior or preanal appendages) forming small, circular, elevated warts; shifted distad, basally surrounded by rim of apicoventral setose lobes. Coxopodites slightly exceeding apex of segment X; forming slightly sinuous rod with dilating apex in lateral view ( Fig. 341); weakly curving mesad apically in ventral view ( Fig. 343). Harpagones sinuate in lateral view ( Fig. 341); basal half broad, S-shaped in ventral view ( Fig. 343). Phallotheca with straight dorsum ( Fig. 344). Endophallus long, running through entire phallotheca; ending in narrow tube at gonopore. Chitinized endothecal process elongating ovoid; curving ventrad. Phallotremal sclerites vertically broad in lateral view. Vestigial, membranous, ventral endothecal lobes nearly invisible.

Holotype male: NEPAL: Chitawan District , Royal Chitawan National Park, 1.xi.1985 [J. Louton] - ( NMNH).

Paratypes: same data as holotype - 9 males, 15 females ( NMNH); 2 males, 1 female ( OPC) .

Distribution. Nepal.

Etymology. chitawana , named after the type locality, Chitawan National Park.