Alcidodes gonzoi Lyal, 2000

Lyal, C. H. C. & Curran, L. M., 2000, Seed-feeding beetles of the weevil tribe Mecysolobini (Insecta: Coleoptera: Curculionidae) developing in seeds of trees in the Dipterocarpaceae, Journal of Natural History 34 (9), pp. 1743-1847 : 1813-1815

Alcidodes gonzoi Lyal , n.sp.

Robust insects, with humeral angles not protruding; length 3.56±6.31 mm (xÅ5 l 4.74, m 4.70), PW 5 1.75±3.13 mm (xÅ5 l2.31, m 2.30), HW5 2.13±3.69 mm (xÅ5 l

2.79, m 2.73) (n 5 l41, m 66) (®gures 31, 64, 82).

Head. Rostrum arises from front of head adjacent to about one-third (l) or half-way (m) down eye; curved, less punctate and more slender in female (RL 5 1.31±2.47 mm, xÅ5 1.84 mm; RL/PW 5 0.71±0.91, xÅ5 0.80; RB/RD5 0.71±1.03, xÅ5

0.84; n 5 66) (®gure 205) than in male (RL 5 1.44±2.34, xÅ5 1.77 mm; RL/PW 5

0.72±0.83, xÅ5 0.77; RB/RD 5 0.82±1.04, xÅ5 0.91; n 5 41) (®gure 206).

Thorax. PL / PW, m, 0.62±0.76 (xÅ5 0.68, n 5 66),, l0.55±0.73 (xÅ5 0.66, n 5 41); prothorax lacking a tubercle behind fore coxae. Elytral interstriae with numerous small indistinct granules, especially proximally and on disc, strial punctures narrower than interstriae, humeri with interstria 9 not projecting beyond 8 in dorsal view

(®gures 31, 49); EL/HW, m, 1.05±1.71 (xÅ5 1.18, n 5 66),, l1.05±1.73 (xÅ5 1.17, n 5 41). Elytral colouration uniformly dark; elytral scales hair-like, uniform. Fore femur with ventral tooth as in ®gures 139, 166, 167, proximal lobe lying along femur and sometimes di cult to see, central lobe at right angles to long axis of femur, distal margin with one or two small teeth but lobe not well developed; fore tibia with ventral tooth, mid and hind tibiae lacking ventral tooth. Ventral margin of metasternum convex, not strongly divergent from elytral margin (cf. ®gure 13).

Abdomen. Male ventrite 5 with very weak median depression (®gure 112); female ventrite 5 with median depression weak (®gure 113) or absent.

Male terminalia. (®gures 359±366). Tergite 7 (®gure 359) with posterior margin weakly concave; one or two plectral tubercles near posterior margin on each side. Segment 8 as ®gure 360. Tegmen (®gures 362±363) with parameres united at base, approximately as long as width of tegminal ring. Membrane between tegmen and aedeagus developed into two inconspicuous lobes (®gure 364). Aedeagus (®gure 364±366) with sides very weakly convex, subparallel, apex rounded; apex with patch of setae on either side of mid-line; dorsal surface of aedeagus sclerotized between ostium and base. Ostiolar sclerites fused to dorsal sclerotization of aedeagus, extending inside endophallus as indistinct elongate sclerites (®gure 365).

Female terminalia. (®gures 367±369) Tergite 7 with 2±3 pairs of plectral tubercles near wing-binding patches (®gure 367). Tergite 8 with broadly rounded posterior margin, disc and median anterior margin more weakly sclerotized than other margins (®gure 368). Spiculum ventrale (®gure 369) with broad part broader than long, not as long as apodeme. Female genitalia as in ®gures 370±371; hemisternites short and broadened towards the base; spermathecal duct arising from bursa distad of its junction with common oviduct; common oviduct on membranous lobe arising from bursa; spermatheca with gland lobe bulbous medially (®gure 370).

Distinguishing features. A. gonzoi is found on the same hosts as ®ve other species of similar size, A. confusus , A. walliorum and A. fugitus (in the crassus group) and A. hopeae and A. sterryorum (both dipterocarpi group). The distinguishing features of A. confusus , A. walliorum and A. fugitus have been discussed above. A. gonzoi may be distinguished from A. hopeae by its much weaker humeral angles (®gures 31, 30, 49, 47). A. sterryorum can be distinguished from both A. hopeae and A. gonzoi by the pro®le of the head (®gures 203±208), the rostrum arising in a more dorsal position in that species than in the other two.

Comments. This species has only been reared from the seeds of Hopea species. Toy’s (1988) record of A. hopeae from H. latifolia Sym. refers to A. gonzoi .

Etymology. The species is named for the muppet Gonzo, with which it shares a characteristic pro®le.

Specimens examined

HOLOTYPE ,, l Malaysia, with labels:`Holo- / type’ and`’ l and`Reared from seed / of Hopea dryobalanoide s ’ and` R. Toy Coll. / B.M. 1992 ±170’ and`87/1’ and (on obverse)`Coll: / FRI’ and` MALAYSIA: / Negri Sembalan, / Pasoh Forest / Reserve / February 1986 #87’ and ` HOLOTYPE / Alcidodes / gonzoi / Lyal 1997’. ( BMNH)

PARATYPES: Malaysia: Peninsular Malaysia : 3 ll, 2mm, same locality and host as holotype ( BMNH); 4 ll, 3mm, Kepong Dipterocarp Arboretum (Toy), reared from seeds of Hopea latifolia (BMNH) .

Indonesia: Borneo: Kalimantan, Gunung Palung Nature Reserve, Cabang Panti Research Station, January± April 1991 (Curran), reared from seeds of: Hopea dyeri (12 ll, 21mm); Hopea ferruginea (17 ll, 29mm); Hopea mengerawan (8 ll, 13mm); Hopea sangal (1 l) ( BMNH, MZB, UMMZ).

Range. Peninsular Malaysia, Borneo.

Hosts. Hopea dyeri Pierre , H. ferruginea Parijs , H. mengerawan Miq. , H. sangal Korth.