Alcidodes fugitus Lyal, 2000

Lyal, C. H. C. & Curran, L. M., 2000, Seed-feeding beetles of the weevil tribe Mecysolobini (Insecta: Coleoptera: Curculionidae) developing in seeds of trees in the Dipterocarpaceae, Journal of Natural History 34 (9), pp. 1743-1847 : 1809-1811

Alcidodes fugitus Lyal , n.sp.

Robust insects, length 4.55±6.45 mm (xÅ5 l5.53, m 5.46), PW 5 2.3±3.25 mm (xÅ5 l2.88, m 2.84), HW 5 3.05±4.25 mm (xÅ5 l3.81, m 3.74) (n 5 l11, m 12) (®gures 29, 61, 80).

Head. Rostrum weakly curved, slightly longer and generally more slender in female ( RL 5 1.88±2.65 mm, xÅ5 2.29, RL/ PW 5 0.75±0.84, xÅ5 0.81; RB/RD 5 0.72±0.88, xÅ5 0.84; n 5 12) (®gure 201) than in male ( RL 5 1.80±2.38, xÅ5 2.19 mm; RL/ PW5 0.74±0.79, xÅ5 0.76; RB/RD 5 0.88±1.02, xÅ5 0.94; n 5 11) (®gure 202); arising near top of eye in both sexes, although higher in male than in female .

Thorax. PL / PW 5 0.66±0.71 (xÅ5 0.68, n 5 23) (no signi®cant sexual diOEerence); prothorax generally swollen behind fore coxae, although tubercle not seen. Elytra with interstriae smooth or weakly wrinkled proximally, strial punctures deep, broad, at least proximally on striae 4 and 5 wider than and partially obscuring interstriae, humeri with interstria 9 projecting beyond 8 in dorsal view, sometimes partially visible in proximal half of length (®gures 29, 44); EL/HW5 1.01±1.13 (xÅ5 1.07, n 5 23), no signi®cant diOEerence between sexes. Elytra uniformly dark in specimens seen; scales fairly uniform in size and shape. Fore femur with ventral tooth subtriangular, lacking proximal lobe, proximal margin at c.90ss to long axis of femur and at least as high as the length of the tooth, proximal and distal margins approximately same length, distal margin dentate or serrate and with relatively steep slope (®gures 137, 164, 165); fore tibia with ventral tooth, mid and hind tibiae lacking ventral tooth or with tooth very small and indistinct. Metasternum strongly produced posteroventrally before hind coxae, its ventral margin strongly diverging from elytral margin .

Abdomen. Ventrite 5 with two caudal setal tufts on prominences on raised posterior margin in both sexes, those of male larger than those of female (®gures 108, 109).

Male terminalia. Tergite 7 as in ®gure 334; one or two plectral tubercles on each side, obscure. Segment 8 as ®gure 335. Genitalia as in ®gures 336±340. Tegmen with parameres fused at base (®gure 336). Aedeagus with apex rounded, downturned, sides weakly concave (®gures 338, 339), distance from ostium to apex less than width of ostium; ostiolar sclerites not fused to aedeagal body (®gure 340).

Female terminalia. (®gures 341±343) Tergite 7 with diagonal row of plectral tubercles on each side of midline (®gure 341). Tergite 8 with broadly rounded posterior margin; disc and anterior margin more weakly sclerotized than rest of tergite (®gure 342). Spiculum ventrale (®gure 343) with broad apical part quadrate, shorter than apodeme. Female genitalia as in ®gures 344±345; hemisternites short, styli long, broad. Spermathecal duct arising from bursa away from junction with common oviduct; spermatheca with gland lobe simple, straight, tubular, striate.

Distinguishing features. The distinguishing features of this species have been discussed above, under A. walliorum .

Comments. The unique specimen reared from the seeds of Hopea ferrugineus is probably truly recorded from this host, given the host range of the closely-related Alcidodes confusus .

Etymology. Specimens of this species were originally mixed with those of other species from the hosts, from which they were separated only with some di culty, hence the concept of them ¯eeing from identi®cation.

Specimens examined

HOLOTYPE, l Indonesia: Kalimantan, with labels:`Holo- / type’ and` INDONESIA: / W. KALIMANTAN; / Gunung Palung / Nat. Pk., Cabang / Panti Res. Sta.’ and`1ss15¾S.110ss05¾E / i± iv.1991 / Lisa M. Curran’ and`LAB. REARED: / Ex: seeds / Hopea / ferrugineus / Dipterocarpaceae ) ’ and`91 Hfer 4’ and` HOLOTYPE / Alcidodes fugitus / Lyal / Lyal det 1996.’ and`652’ ( MZB).

PARATYPES: 10 ll, 12mm, same locality as holotype, but host Hopea sangal (MZB, BMNH, UMMZ) .

Range. Borneo.

Hosts. Hopea ferruginea Parijs , H. sangal Korth.