Craspedophorus glaber Bates, 1886

Häckel, Martin, 2017, A contribution to the knowledge of the subfamily Panagaeinae Hope, 1838 from Africa. Part 3. Revision of the Craspedophorus strachani and C. brevicollis groups (Coleoptera: Carabidae), Zootaxa 4330 (1), pp. 1-67 : 12-13

6. Craspedophorus glaber Bates, 1886

( Plate 7 , Figs 61a,b, 62 a,b; Plate 8 , Fig. 68a, Plate 9 , Fig. 61c, Plate 13 , Fig. 105)

Craspedophorus glaber Bates, 1886a: 196 (type locality “Mount Cameroons [=Cameroon, Southwest]”). Lorenz 2005: 320. Häckel and Farkač 2012: 81.

Type material. “Holotype (♀): “Cameroons [handwritten in black on white label]// Craspedophorus / glaber / Chaud. [handwritten in black on white label]// Ex-Musaeo / H. W. Bates / 1892 [printed in black on white label]// Type [printed] / glaber [handwritten in black on red label]” ( Plate 7 , Fig. 62a, MNHN, Oberthür/Bates Collection). Paratypes. 2♂, 2♀: “Cameroons [handwritten in black on white label]// Ex-Musaeo / H. W. Bates / 1892 [printed in black on white label]// Paratype [printed in black on red label]” (MNHN, Oberthür/Bates Collection), 1♂ same labeled as holotype except “Museum Paris / Coll. Ch. Alluaud [printed in black on blue label]// glaber / Bates / ex typis. [handwritten in black on white label]” (MNHN). 1♀ erroneously labeled and pinned to series with paratypes of C. lafertei Murray : “Paratype [printed in black]/ Lafertei Murr. [handwritten in black on red upper pinned label]/ / Old Calabar [printed in black on white lower pinned label]// pinned first from left in the series labeled: “ Lafertéi? / Murray / Afrique occident. / Sallé [handwritten in black on white box label in Chaudoir's Collection]” ( Plate 7 , Fig. 62b, MNHN).

Additional material examined. Cameroon. 1♀: “ Kamerun / H. Rolle, Berlin W. // Cr. glaber Bat. / P. Basilewsky det., 1954” ( MNHN) . Centre. 1♂ labeled as “Comp. typ.” by Basilewsky 1951: “Yaoundé” ( Plate 7 , Fig. 61b, MRAC). Southwest. 1♂, 1♀: “ Mungo [River] Mukonje Farm ” ( MRAC) . 1♂: “SWpr. 3.5 km ne Bakingili, N: 4°5.431'; E: 9°3.131', 400 m ” ( Plate 7 , Fig. 61a, Plate 9 , Fig. 61c, Plate 13 , Fig.105, cMH).

Note. This species is based on number of specimens labeled “ Cameroons ” collected in Cameroon Mountains. Description (in part, see Bates 1886a: 196). “Length 16 mm. Body short, wide, parallel-sided, black, glossy, oral parts, antennae (except basal articles), genae, tibiae and tarsi ferruginous, each elytron with two large yellow maculae, humeral macula reaches from V to IX interval, preapical one reaches from IV to VIII interval; head wide, eyes strongly convex; pronotum similar to that in C. grossus [ Hope, 1842], but with lateral margins more widely elevated and narrowed posteriorly; elytra short, almost ovoid, striae deeply impressed and punctured, intervals fully smooth, without punctuation“ [from Latin]. “In everything except the more triangular apical joint of the palpi this species accords with the first sub-section of Chaudoir's genus, Craspedophorus (1878: 11) . It has very short mesothoracic episterna, much broader than long, strongly crenulated anterior edges of ventral segments, and in the form of the thorax resembles closely C. grossus [ Hope, 1842]. It differs, however, from C. grossus and from C. ruficrus [LaFerté-Sénectere, 1851] and C. lafertei [ Murray, 1857] in the more prominent eyes and narrower neck (the crown very coarsely punctured). It is a much shorter and less convex species than G. grossus . The thorax is relatively broader and more angularly dilated in the middle, and its surface is very strongly punctured, the punctures in the middle being distinctly separated from each other. In the elytral surface being quite destitute of hairs and the interstices impunctate it is distinguished from all allied species ( Plate 9 , Fig. 61c). The anterior red spot of the elytra is quadrate, and reaches nearly to the base, the sub-apical spot is shorter and more transverse, but variable in length. According to the shape of the apical joint of the palpi the species would belong to Epicosmus according to Chaudoir, but all the other characters are those of Craspedophorus ”. Bates correctly places this species in Craspedophorus (here C. strachani group) in spite of his objections regarding the shape of terminal labial palpi. Those being examined in males (not in females or small males), the prolongated kidney-form shape of the terminal palpi is obvious, thus the placement in the genus (according to Chaudoir) and in the newly-defined C. strachani group seems to be correct.

Ditribution. Central, southwestern Cameroon; Gabon, south-eastern Nigeria: Cross River State.

Figures

PLATE 7. Habitus of Craspedophorus strachani group VI (Scale bar: 10 mm). 56. C. phenacoides n. sp., HT (male); 57. C. phenacoides n. sp., PT female, Cameroon, Centre; 58. C. lemariei n. sp., HT (male); 59 a. C. ruficrus (Chaudoir, 1861) HT (female); b. C. ruficrus (Chaudoir, 1861), female, Guinea Equatorial; 60. C. dicranulothorax n. sp., HT (female); 61 a. C. glaber Bates, 1886, male, Cameroon, Southwest; b. C. glaber Bates, 1886, male, Cameroon, Centre (comparative type Basilewsky 1954); 62 a. C. glaber Bates, 1886, HT (female); b. C. glaber Bates, 1886, female, “ Old Calabar ” (labeled as C. lafertei, MNHN); 63. C. lafertei Murray, 1857, lectotype female, established by Basilewsky 1960; 64. C. lafertei Murray, 1857, female, “ W Africa ” (paralectotype established by Chaudoir 1879, MNHN); 65. C. dicranothorax (Alluaud, 1915), HT (male); 66. C. dicranothorax (Alluaud, 1915), female, Central Africa.

PLATE 8. Craspedophorus strachani group, C. ruficrus complex, head and pronotum. 67 a. C. ruficrus (Chaudoir, 1861), female Guinea Equatorial. 68 a. C. lemariei n. sp., HT (male). 69 a. C. dicranulothorax, HT (female). 70. C. phenacoides n. sp., HT (male). 71. C. r. ruficroides n. sp., HT (male).

PLATE 9. Craspedophorus strachani group, C. ruficrus complex. Elytra (left humerus from oblique view): Fig. 61 c. C. glaber Bates, 1886, male, Cameroon; Fig. 67 b. C. ruficrus (Chaudoir, 1861) female Guinea Equatorial. Fig. 68 b. C. lemariei n. sp. HT (male); Fig. 69 b; C. dicranulothorax n. sp., HT (female). Fig. 72. Pronotum in different species of the C. strachani group (design), a-c. C. strachani s. str. subgroup, d, e) C. muata subgroup.

PLATE 13 Craspedophorus strachani group II, aedeagus in frontal and right lateral views: 101. C. chevalieri (Alluaud, 1915), Central Africa; 102. C. cuneatus cuneatus (Alluaud, 1915), Central Africa; 103. C. cuneatus paromius Basilewsky, 1987, Orientale, DR Congo; 104. C. cuneatus rotundatus new subspecies, HT; 105. C. glaber Bates, 1886, Cameroon, Southwestern; 106. C. ruficroides ruficroides new species, HT; 107. C. ruficroides thomsoni new subspecies, PT RCA; 108. C. phenax Basilewsky, 1987, PT Fontem, Cameroon, West; 109. C. congoanus Kolbe, 1889, Eala Bokatola-Bikoro, Équateur, DR Congo; 110. C. phenacoides new species, HT; 111. C. lemariei new species, HT.